ON :
Calinaga funeralis Monastyrskii & Devyatkin, 2000 OD : Atalanta 31 (3/4):477-478, pl.18, figs.5(HT,♂), 6(HT,♂Un). TL : Ba Be, Bac Can, N.Vietnam. Distribution : N.Vietnam. |
Calinaga lhatso coniaensis Funahashi, 2003 : Wallace 8:9-10, pl.3, fig.13(HT,♂). TL. Mt. Pia Oac, N.Vietnam.
funeralis :
Holotype ♂, 02.Apr.1997. Ba Be N.P., Bac Can prov., N.Vietnam. A.L.Monastyrskii leg. (BNHN.)
Paratypes : 5♂, same locality as the holotype. (NHML & MNHN.)
coniaensis :
Holotype ♂, Apr.1995. Mt. Pia Oac, alt.1911m, N.Vietnam. (in coll. A.Funahashi)
Paratypes : 21♂ 8♀, same locality as the holotype. (in coll. A.Funahashi ; Morita)
[ VIETNAM ]
Calinaga funeralis ; Monastyrskii & Devyatkin,2015 : 45. (N: Bac Can ; Lao Cai)
Calinaga funeralis ; Monastyrskii,2019 : 4(1):45, pl.1, figs.5a(HT,♂), 5b(HT,♂Un). (N: Ba Be; Dong Phuc; Pia Oac)
Calinaga funeralis funeralis ; Tshikolovets,2020 : 85-87, pl.XXXIX, figs.1(HT of funeralis,♂),2(HT of funeralis,♂Un),3(PT of funeralis,♂),4(PT of funeralis,♂Un),5(PT of funeralis,♂),6(PT of funeralis,♂Un),7(HT of coniaensis,♂),8(HT of coniaensis,♂Un). (N.Vietnam: Bac Can; Mt. Pia Oac)
Tshikolovets (2020: 79) treated lhatso, senseiensis, pacifica, funebris, funeralis and dubernardi as "superspecies lhatso" and these taxa as separate species.
Todisco et al. (2024) treat lhatso and funebris as subspecies of lhatso based on molecular analysis,
and funeralis, senseiensis, dubernardi and yaonica as incertae sedis (? subspecies of lhatso).
Furthermore, the taxon pacifica is treated as incertae sedis (possibly a separate species).
Very interestingly, their phylogenetic analysis shows that funeralis and sensiensis do not belong to the lhatso group but in the brahma group.
It is possible that they belong to a different species group than their external features, based on morphological similarity due to their being mimetic species.
I am tentatively treating funeralis as a distinct species until it is determined which group it belongs to.
The genus Calinaga is extremely difficult to identify by morphological characteristics and requires a comprehensive verification, including examination of genitalia and further DNA analysis.
Monastyrskii, A.L. & Devyatkin, A.L.,2000 : New taxa and new records of butterflies from Vietnam (Lepidoptera, Rhopalocera).
Atalanta 31(3/4):471-492, col.pls.18-21a
Funahashi, A.,2003 : Description of a new subspecies of Calinaga lhatsuo Oberthür, 1893 from Mt.PiaOac, N.Vietnam.
in Butterflies of Vietnam part.2., Descriptions of 19 New subspecies of Butterflies from Vietnam.
Wallace 8:9-10, pl.3.
Tshikolovets, V.,2020 : The Genus Calinaga Moore, [1858] (Lepidoptera: Nymphalidae, Calinaginae).
Tshikolovets Publications. 159 pp, 56 pls.
Todisco, V., Basu, D.N., Prosser, S.W.J., Russell, S., Mutanen, M., Zilli, A., Huertas, B., Kunte, K. & Vane-Wright, R.,2024 :
DNA barcodes from over-a-century-old type specimens shed light on the taxonomy of a group of rare butterflies (Lepidoptera: Nymphalidae: Calinaginae).
PLoS ONE 19(7): e0305825. "https://doi.org/10.1371/journal.pone.0305825"
2020.01.31 - 2024.10.31. |