Papilio helenus helenus   Linnaeus,1758

Papilio helenus helenus ♂ Up. Papilio helenus helenus ♂ Un. Papilio helenus helenus ♀ Up. Papilio helenus helenus ♀ Un.
♂, 1995.03.17. Umphang, Tak, Thailand. ♀, 1987.03.30. Doi Suthep, Chiang Mai, Thailand.
Papilio helenus helenus ♂ Up. Papilio helenus helenus ♂ Un. Papilio helenus helenus ♀ Up. Papilio helenus helenus ♀ Un.
♂, 1989.08.24. Doi Pee Ban Nam, Xaignabouri, Laos. ♀, 1991.10.19. Doi Phu Kha, Nan, Thailand.

ON : Papilio helenus  Linnaeus,1758
OD : Syst.nat. (ed.10) 1:459.
TL : "Asia".



Distribution :
Nepal, India to Myanamr, Thailand, Laos, Cambodia, 
Vietnam, Hainan, S.China, Langkawi, W.Malaysia, Tioman.
Papilio helenus helenus Map

Synonyms

Papilio helenus ♂ab.rufatus  Rothschild,1895 : Novit. zool. 2(3):286.  TL. India ; Burma ; Malacca ; China.
Papilio helenus aulus  Fruhstorfer,1908 : Ent. Wochenbl. 25:38.  TL. Hainan. (BMNH)
Papilio helenus ab.aurea  Boullet & Le Cerf,1912 : Bull. Soc. ent. Fr. 1912(11):247.  TL. Pnom-Penh, Cambodia. (MNHN)
Papilio helenus aspadantus  Fruhstorfer,[1916] : Arch. Naturgesch. (A)81(11):75.  TL. Malayische Halbinsel. (BMNH)


Type Materials Information

aulus ; ♂, Type (red), #149316, Hainan, Whitehead. (BMNH. examined.)
aspadantus ; ♂, Type (red), #149317, Malaka, H.Fruhstorfer (BMNH. examined.)
aurea ; Syntype 1♂, Pnom-Penh, Cambodge. (MNHN.)


Records taken from Literature

[ THAILAND ]
Papilio helenus helenus ; Godfrey,1916 : 110.
Papilio helenus helenus ; Pendlebury,1923 : 23-24. (Nakhon Sri Thammarat: Khao Luang ; Tai Sai River)
Papilio helenus helenus ; Godfrey,1927 : 120. (North)
Papilio helenus helenus ; Godfrey,1930 : 212. (Prae / Muak Lek , East / Chonburi: Sriracha / Chiang Rai / Nakhon Sri Thammarat)
Papilio helenus helenus ; Godfrey,1932a : 264. (Peninsula)
Papilio helenus helenus ; Reeves,1966 : 10. (Nakhon Nayok: Khao Yai)
Papilio helenus helenus ; Reeves,1967a : 120. (Nakhon Nayok: Khao Yai)
Papilio helenus helenus ; Pinratana,1977 : 36, fig.P25,♂,♂(Un). (all over)
Papilio helenus helenus ; Pinratana,1992 : 22,42 pl.28, fig.25,♂,♀, pl.29, fig.25,♂(Un),♂(ab.Un). (Chiang Mai / Chaiyaphum / Chanthaburi / Chonburi / Kanchanaburi / Nakhon Nayok / Ratchaburi / Ranong)
Papilio helenus helenus ; Katsuyama et al.,2006 : 46, fig.27♂. (Nan: Ban Nam Poon)
Papilio helenus helenus ; Ek-Amnuay,[2007] : 78, pl.10, fig.P26,♂,♀. (Kanchanaburi)
Papilio helenus helenus ; Les Day,2011- : Samui Butterflies, fig.♂,♂[♀](Un). (Koh Samui)
Papilio helenus helenus ; Kimura et al.,2011 : 123, fig.♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok / Doi Chiang Dao / Doi Suthep / Doi Inthanon / Thung Saleng Luang / Nam Nao / Phu Khiewo / Khao Yai / Nakohn Nayork / Wan Nam Khiewo / Kao Soi Dao / Kratchin WF / Khao Khiewo / Doi Musso / Sai York Noi / Hua Hin / Muang Shone / Khlong Naka / Ka Chong / Than To WF)
Papilio helenus helenus ; Ek-Amnuay,2012 : 78, pl.10, fig.P24,♂,♀. (Kanchanaburi)

[ LAOS ]
Papilio helenus helenus ; Motono & Negishi,1989 : 46 pl.5 figs.5♂,6♂(Un),7♀,8♀(Un). (everywhere)
Papilio helenus helenus ; Osada, Uémura & Uehara,1999 : 200, pl.11, figs.♂(dsf),♂(dsf,Un),♀(dsf). (Lak Sao / Nong Het)
Papilio helenus helenus ; Nimura & Wakahara,2005 : 38. (Thabok / Nakai)
Papilio helenus helenus ; Cotton & Racheli,[2007] : 349-350. (throughout Laos)
Papilio helenus helenus ; Onodera,2009b : 36-37. (Xieng Kouang: Nam Noen)
Papilio helenus helenus ; Nakamura & Wakahara,2012 ; 41.

[ CAMBODIA ]
Papilio helenus ab.aurea ; Dubois & Vitalis de Salvaza,1914 : 149. (Cambodge)
Papilio helenus ab.aurea ; Dubois & Vitalis de Salvaza,1921 : 12. (Cambodge)
Papilio helenus helenus ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)

[ VIETNAM ]
Papilio helenus ; Joannis,1901 : 333. (Tonkin: Cao Bang)
Papilio helenus ; Fruhstorfer,[1903] : 173. (Tonkin: Chiem-Hoa / Annam: Phuc-Son; Xom-Gom)
Papilio helenus helenus ; Jordan,1908-1909 : 54. (Tonkin)
Papilio helenus ; Lemee,1950 : 4. (Hagiang / Hanoi)
Papilio helenus helenus ; Metaye,1957 : 82 (North / Central / South)
Papilio helenus daksha ; Metaye,1957 : 82 (North / Central / South)
Papilio helenus helenus ; Ikeda et al.,1998 : 17, figs.3:15♂,16♂(Un),4:1♀,9:6(♂genitalia). (Cuc Phuong)
Papilio helenus helenus ; Shinkai,1999 : 53. (North: Tam Dao / South: Bao Lac)
Papilio helenus helenus ; Miyazaki, Saito & Saito,2006 : 7, figs.Pa-13♂,♀. (Lam Dong: Dambri ; Gung Re ; Madagui)
Papilio helenus ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Papilio helenus helenus ; Monastyrskii,2007 : 55-56, pl.13, figs.2a♂,2b♂(Un),2c♀. (throughout Vietnam)


Specimens Examined

[ THAILAND ]
Mae Hong Son, Ban Ten Doi.
1992.01.30
1♂


R.Phongpiyamit (YI)
Mae Hong Son, Ban Huai Pung.
1993.02.03
1♂


P.Sukkit (YI)
Chiang Mai, Chiang Dao.
2014.02.15
1♂


Y.Inayoshi (photo)
Chiang Mai, Doi Suthep.
1983.03.04
3♂


Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1983.03.05
2♂


Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1987.03.30


1♀
Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1983.04.02
1♂


Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1984.04.14


1♀
Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1985.08.16
1♂


Y.Shiraishi (YI)
Lampang, Mae Mo.
1993.10.25


1♀
M.Nishimura (MN)
Nan, Doi Phu Kha.
1992.01.10
1♂


Local Catcher (YI)
Nan, Doi Phu Kha.
1992.02.25


1♀
Local Catcher (YI)
Nan, Doi Phu Kha.
1991.03.12


1♀
Y.Inayoshi (YI)
Nan, Doi Phu Kha.
1992.03.13
1♂


Local Catcher (YI)
Nan, Doi Phu Kha.
1991.06.--
2♂


Local Catcher (YI)
Nan, Doi Phu Kha.
1991.10.19


1♀
Local Catcher (YI)
Tak, Umphang.
1995.03.17
1♂


M.Nishimura (MN)
Chanthaburi, Khao Soi Dao.
1984.03.07
3♂


Y.Inayoshi (YI)
Chanthaburi, Khao Soi Dao.
1983.03.27
1♂


Y.Inayoshi (YI)
Trat, Koh Chang.
1996.03.13
1♂


Y.Inayoshi (YI)
Ranong, Bo Nam Ron.
1983.03.22
1♂


Y.Inayoshi (YI)
Ranong, Muang Chone.
1984.03.27
1♂


Y.Inayoshi (YI)
Ranong, Muang Chone.
1987.04.15
1♂


Y.Inayoshi (YI)
Phuket, Tone Sai WF.
1986.11.05
1♂


A.M.Cotton (YI)
Nakhon Si Thammarat.
2001.04.12
2♂


P.Sukkit (PS)

[ LAOS ]
Xaignabouri, Doi Pee Ban Nam.
1989.08.21
1♂


P.Sukkit (YI)
Xaignabouri, Doi Pee Ban Nam.
1989.08.23
2♂


P.Sukkit (YI)
Xaignabouri, Doi Pee Ban Nam.
1989.08.24
2♂


P.Sukkit (YI)
Xaignabouri, Doi Pee Ban Nam.
1989.08.30
1♂


P.Sukkit (YI)
Xaignabouri, Doi Pee Ban Nam.
1989.08.31
1♂


P.Sukkit (YI)
Lak Sao, Nam Phao.
2005.02.26-03.11
3♂
1♀
A.M.Cotton (AC)
Lak Sao, Nam Phao.
2005.03.21-03.30
1♂


A.M.Cotton (AC)
Lak Sao, Nam Phao.
2005.06.19
1♂


A.M.Cotton (AC)
Lak Sao, Nam Phao.
2005.06.21
1♂


A.M.Cotton (AC)
Lak Sao, Nam Phao.
2005.07.29
1♂


A.M.Cotton (AC)
Thabok, Tad Leuk.
2005.02.26-03.11
26♂


A.M.Cotton (AC)
Thabok, Tad Leuk.
2005.03.21-03.30
2♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.02.26-03.11
2♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.03.21-03.30
8♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.06.14
6♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.06.23
6♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.08.08-08.10
31♂
1♀
A.M.Cotton (AC)
Hinboun, Koun Kham.
2005.02.26-03.11
2♂


A.M.Cotton (AC)
Hinboun, Koun Kham.
2005.03.21-03.30
3♂


A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.06.17
2♂


A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.07.30-07.31
1♂
1♀
A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.08.01-08.07
11♂
2♀
A.M.Cotton (AC)
Hinboun, Phu Pha Man.
2005.08.05-08.07
1♂
3♀
A.M.Cotton (AC)
Cham Pasak, Paksong.
1996.03.25
1♂


Y.Inayoshi (YI)

[ VIETNAM ]
Lam Dong, Dambri.
2004.01.01
1♂


K.Saito (KS)
Lam Dong, Dambri.
2003.09.15-09.20
4♂
1♀
K.Saito (KS)
Lam Dong, Dambri.
2002.12.30
1♂


K.Saito (KS)
Lam Dong, Dai Duc Me.
2003.12.29
1♂


K.Saito (KS)
Lam Dong, Madagui.
2006.10.03
1♂


K.Saito (KS)
Lam Dong, Loc Tan.
2006.10.04
1♂


K.Saito (KS)


Notes
Literature

Linnaeus, C.,1758 : Systema naturae per regna tria naturae,secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis.
Holmiae, Laurentil Salvii. (Edition 10). 1(Animalia):[iv]+824pp.

Rothschild, L.W.,1895 : A revision of the Papilios of the Eastern Hemisphere, exclusive of Africa.
Novit. zool. 2(3):167-463, pl.4.

Fruhstorfer, H.,1908 : Neue ostasiatische Rhopaloceren.
Ent. Wochenbl. 25:38.

Boullet, E. & F. Le Cerf,1912 : Descriptions sommarires de formes nouvelles de Papilionidae (Lep).
de la collection du Museum de Paris (2e note).
Bull. Soc. ent. Fr. 1912(11):246-247.

Fruhstorfer, H.,[1916] : Rhopaloceren aus Hollandisch-Neu-Guinea.
Arch. Naturgesch. (A)81(11):61-78.

1996.10.02 - 2014.06.14.