Heliophorus ila nolus  Eliot,1963

Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ds), Chiang Mai, Thailand. ♀ (ds), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ws), Chiang Mai, Thailand. ♀ (ds), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ws), Tak, Thailand. ♀ (ws), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un.
♂ (ds), Chiang Mai, Thailand. ♂ (ds), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia
♂ genitalia, Kanchanaburi, Thailand. ♂ genitalia (transition form), Nan, Thailand.

ssp.nolus (transition populations to ssp.urius)
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ds), Lampang, Thailand. ♀ (ds), Lampang, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ds), Chiang Mai (Doi Saket dist.), Thailand. ♀ (ds), Nan, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ds), Nan, Thailand. ♀ (ws), Nan, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♀ Up. Heliophorus ila nolus ♀ Un.
♂ (ws), Nan, Thailand. ♀ (ws), Nan, Thailand.
Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un. Heliophorus ila nolus ♂ Up. Heliophorus ila nolus ♂ Un.
♂ (ds), Xieng Khouang, Laos. (photo by H.Onodera) ♂ (ws), Xieng Khouang, Laos. (photo by H.Onodera)

ON : Heliophorus ila nolus Eliot,1963
OD : Entomologist 96(1203):175-176.
TL : Taok Plateau. Tenasserim, Burma. (♂, NHML)
Distribution : Myanmar, Thailand. N. to C.Laos. [N.Vietnam]

● ssp.nolus
ssp.nolus (transition zone to ssp.urius)
ssp.urius
Heliophorus ila map

Type Materials Information

Holotype ♂, Taok Plateau, 3000 ft., Tenasserim, Burma. 1924.i.1. C.H.Stockley. (NHML)


Records taken from Literature

[ THAILAND ]
Heliophorus epicles indicus ; Shirozu & Saigusa,1962 : 60-61, pl.9, figs.67,♂,68,♂(Un), Textfig.27,♂genitalia. (Doi Inthanon)
Heliophorus ila nolus ; Pinratana,1981 : 72, pl.11, figs.21,♂,22,♀,23,♂(Un). (Chiang Mai)
Heliophorus ila nolus ; Ek-Amnuay,[2007] : 592, pl.268, fig.L167,♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok; Mae Hong Son; Doi Phu Kha)
Heliophorus ila nolus ; Okamoto et.al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
"Heliophorus brahma magoka ; Ek-Amnuay,2012 : 618, pl.280, fig.L186,♀. (in part)"
Heliophorus epicles latilimbata ; Ek-Amnuay,2012 : 618, pl.280, figs.L189,♀,♀(Un). (in part)
Heliophorus ila nolus ; Ek-Amnuay,2012 : 618, pl.280, figs.L191,♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok; Mae Hong Son; Doi Phu Kha)
Heliophorus epicles latilimbata ; Kimura et al.,2014 : 151, figs.♀,♀(Un). (Doi Pha Hom Pok) (in part)
Heliophorus ila nolus ; Kimura et al.,2014 : 151, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Pha Hom Pok; Chiang Dao; Doi Pui; Doi Suthep; Doi Inthanon; Maethro; Khun Lao; Phu Kradung; Nam Nao)
Heliophorus epicles latilimbata ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Kamphaeng Phet) (accessed: 2021.09.03)
Heliophorus ila nolus ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Chiang Mai) (accessed: 2021.09.03)

[ VIETNAM ]
Heliophorus ila nolus ; Yago et al.,2005 : 366, 369, figs.6,♂,♂(Un). (Dong Van; Muong Nhe; Hoang Lien)
Heliophorus ila nolus ; Monastyrskii & Devyatkin,2015 : 55. (N: Ha Giang; Lao Cai; Lai Chau)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 10♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai (Fang dist; Chiang Dao dist; Chiang Mai dist; Mae Chaem dist; Omkoi dist.) : 25♂ 16♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Kanchanaburi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

--------------------------------------------------
transition between ssp.nolus and ssp.urius

Chiang Rai (Thoeng dist.) : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai (Doi Saket dist; Mae On dist.) : 12♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 10♂ 3♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phisanulok : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 16♂ 11♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Luang Nam Tha : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Luang Phabang : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Houaphan : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Xieng Khouang : 3♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
The butterfly is common in montane forest at moderate to high elevations (alt.700m - 1800m). Frequently, it occurs sympatric with epicles at 700m-1100m and with indicus and brahma at 1600m-1800m.
This species and indicus of this region are very similar, and it is difficult to identify them by markings. Examination of genitalia is necessary for accurate identification.
The populations in Indochina are highly variable in terms of individual, seasonal and geographical variation in both markings and genitalia. Frequently, male individuals with orange patch on the UpF are found. The populations in W.Thailand, Tak and Kanchanaburi, are definitely ssp.nolus. The characteristics of genitalia are costal shoulder ill-defined, transtilla aborted and vanni long with truncate tips. In the area from W.Thailand to the mountain range in west of Chiang Mai city in N.Thailand is generally nolus, however occasionally individuals transitioning to urius are seen. On the other hand, from the eastern mountain range of Chiang Mai city to north part of Laos, it is generally a transition form to urius, but occasionally nolus form are found. In the genitalia of this transitional individuals, the transitilla is slightly more developed, the vanni are shorter. In terms of markings, post-discal black spots on the UnF tend to be slightly more prominent. (especially in the rainy season). In Xieng Khouang, Laos, There are both intermediate form which is more closer to urius than nolus and ssp.urius occur sympatrically. Yago et al.,2005 and Monastyrskii & Deviatkin,2015 recorded nolus in N.Vietnam. I have tentatively mapped them as nolus, however it seems more likely they are a transitional population. Further taxonomic study is needed.

Male genitalia of each populations
ssp.nolus :
Kanchanaburi, Thailand.
Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia
ssp.nolus (intermediate form) :
Loei, Thailand.
Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia Heliophorus ila nolus ♂ genitalia
ssp.nolus (intermediate form) :
Xieng Khouang, Laos.
(photo by J.Uehara & H.Onodera)
Heliophorus ila urius ♂ genitalia Heliophorus ila urius ♂ genitalia Heliophorus ila urius ♂ genitalia
ssp.urius :
Bolikhamxai, Laos.
Heliophorus ila urius ♂ genitalia Heliophorus ila urius ♂ genitalia Heliophorus ila urius ♂ genitalia

Literature

Eliot, J.N.,1963 : The Heliophorus epicles (Godart, 1823) (Lepidoptera : Lycaenidae) Complex
Entomologist 96(1203):169-180, 14 figs.

Yago, M., M.Wakabayashi, B.Tanaka. & A.L.Monastyrskii,2005 : Species biodiversity and life histories of the genus Heliophorus (Lepidoptera: Lycaenidae) in Vietnam.
Report on Insect Inventory Project in Tropic Asia (TAIIV)(2005):365-370.

Heliophorus ila nolus ♂ Heliophorus ila nolus ♀
♂ (ds, territorial behavior), Chiang Mai, Thailand. ♀ (ws), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ Heliophorus ila nolus ♀
♂ (ws), Chiang Mai, Thailand. ♀ (ws), Chiang Mai, Thailand.
Heliophorus ila nolus ♂ Heliophorus ila nolus ♂
♂ (ws), Chiang Mai, Thailand. ♂ (ds), Tak, Thailand.
Heliophorus ila nolus ♂ Heliophorus ila nolus ♀
♂ (ws), Lampang, Thailand. ♀ (ws), Lampang, Thailand.
Heliophorus ila nolus ♂ Heliophorus ila nolus ♂
♂ (ds), Lampang, Thailand. ♀ (ds), Lampang, Thailand.
Heliophorus ila nolus ♂ Heliophorus ila nolus ♂
♂ (ds, territorial behavior), Loei, Thailand. ♂ (ws), Nan, Thailand.

1999.07.11 - 2021.09.30.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.