Heliophorus ila urius  Eliot,1963

Heliophorus ila urius ♂ Up. Heliophorus ila urius ♂ Un. Heliophorus ila urius ♂ Up. Heliophorus ila urius ♂ Un.
♂ (ws), Bolikhamxai, Laos. ♂ (ws), Bolikhamxai, Laos.
Heliophorus ila urius ♂ Up. Heliophorus ila urius ♂ Un. Heliophorus ila urius ♂ genitalia Heliophorus ila urius ♂ genitalia
♂ (ws), Xieng Khouang, Laos. (photo by H.Onodera) ♂ genitalia, Lak Sao, Laos.

ON : Heliophorus ila urius Eliot,1963
OD : Entomologist 96(1203):176.
TL : Manhao, S.Yunnan. (♂, NHML)
Distribution : C. to E.Laos, N.to C.Vietnam, S.Yunnan.

● ssp.nolus
ssp.nolus (transition zone to ssp.urius)
Heliophorus ila map

Type Material Information

Holotype ♂, Manhao, S.Yunnan. (NHML)

Records taken from Literature

[ LAOS ]
Heliophorus ila nolus ; Osada, Uémura & Uehara,1999 : 216. pl.112, figs.♂,♂(Un),♀. (Lak Sao)
Heliophorus ila nolus ; Nakamura & Wakahara,2012 : 52.

Heliophorus ila chinensis ; Yago et al.,2005 : 366, 369, figs.5,♂,♂(Un). (Pu Mat)
Heliophorus ila chinensis ; Monastyrskii & Devyatkin,2015 : 55. (C: Nghe An to Lam Dong)

Specimens Examined

[ LAOS ]
Xieng Khouang : 2♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 6♂ (incl.photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Cao Bang : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

The butterfly is not rare in montane forest at moderate to high elevations.
The genitalia of this taxon is very different from ssp.nolus. As mentioned in the section on ssp.nolus, there is a transition zone from N.Thailand to N.Laos where populations with intermediate characteristics to ssp.nolus and ssp.urius are found. Both ssp.urius and intermediate form which is more closer to urius than nolus occur sympatric in Xieng Khouang, Laos. And in Lak Sao, it is only found ssp.urius. Yago et al.,2005 and Monastyrskii & Deviatkin,2015 recorded nolus in N.Vietnam. I have tentatively mapped them as nolus, however it seems more likely they are a transitional population. Further taxonomic study is needed.


Eliot, J.N.,1963 : The Heliophorus epicles (Godart, 1823) (Lepidoptera : Lycaenidae) Complex
Entomologist 96(1203):169-180, 14 figs.

Yago, M., M.Wakabayashi, B.Tanaka. & A.L.Monastyrskii,2005 : Species biodiversity and life histories of the genus Heliophorus (Lepidoptera: Lycaenidae) in Vietnam.
Report on Insect Inventory Project in Tropic Asia (TAIIV)(2005):365-370.


A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.