Phaedyma columella martabana  (Moore,1881)

Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Un. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Un.
♂ (dsf), Chiang Mai, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Un. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Un. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Un.
♂, Mae Hong Son, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Up.
♂ (dsf), Champasak, Laos. ♀, Vientiane, Laos.
Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Up.
♂, Surat Thani (Koh Phangan dist.), Thailand. ♀ (dsf), Phetchaburi, Thailand.
Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♂ Up. Phaedyma columella martabana ♀ Up. Phaedyma columella martabana ♀ Up.
♂ (dsf), Bin Thuan, Vietnam. (photo by K.Saito) ♀, Dong Nai, Vietnam. (photo by K.Saito)

ON : Neptis martabana Moore,1881
OD : Trans. ent. Soc. Lond. 29(3):310.
TL : Rangoon. (♂, NHML)



Distribution : Myanmar, Thailand, Laos, Cambodia, C. & S.Vietnam.
Phaedyma columella martabana Map

Synonym

Phaedyma columella alesia Fruhstorfer,1905 : Ent. Zeit. 19(17):90. TL. Siam ; S.Annam. (♂, MNHN)


Records taken from Literature

[ THAILAND ]
Neptis columella martabana ; Fruhstorfer,1912 : 615. (Siam)
Neptis columella martabana ; Godfrey,1916 : 129. (Chonburi: Don Rek hills / Sriracha)
Neptis columella martabana f.martabna ; Godfrey,1930 : 282. (Chiang Mai / Muak Lek / Chonburi: Sriracha / Bangkok / Chumphon / West: Pan Yao)
Neptis columella martabana f.alesia ; Godfrey,1930 : 282. (N.E.: Nam Pat / W: Pan Yao)
Phaedyma columella martabana ; Eliot,1969 : 120. (Siam)
Phaedyma columella martabana ; Pinratana,1979 : 53, figs.N93a,♂,♂(Un). (Chiang Mai / Chanthaburi / Kanchanaburi / Nakhon Ratchasima / Nakhon Sri Thammarat / Saraburi)
Phaedyma columella singa ; Pinratana,1979 : 53, figs.N93b,♀,♀(Un). (Trang)
Neptis columella martabana ; Pinratana & Eliot,1996 : 54, pl.34, figs.87,♂,♂(Un). (Chiang Mai / Chaiyaphum / Nakhon Ratchasima / Nakhon Nayok / Chanthaburi / Kanchanaburi / Nakhon Sri Thammarat / Trang)
Phaedyma columella martabana ; Yamaguchi & Aoki,2000 : 255, fig.98♂. (Mukdahan: Phu Sa Duak Bua ; Phu Mu)
Phaedyma columella martabana ; Yamaguchi & Aoki,2002 : 28. (Ubon Ratchathani: Ban Na Pho Krang ; Phu Chong Na Yoi ; Nam Tok Soi Sawan ; Kaeng E-Kewo ; Phu Hin Dang / Mukdahan: Ban Lao Kram ; Nam Tok Kham Cha I ; Ban Na Lak ; Phu Huay Sing ; Ban Gok Tum)
Phaedyma columella martabana ; Yamaguchi, Aoki & Akiyama,2005 : 51. (Mukdahan: Ban Lao Kram ; Phu Hin Khan / Ubon Ratchathani: Nam Tok Soi Sawan ; Ban Ba hai ; Phu Chong Na Yoi)
Phaedyma columella martabana ; Katsuyama et al.,2005 : 70, fig.154♂. (Umphang: Umphang ; Tha Sai)
Phaedyma columella martabana ; Katsuyama et al.,2006 : 50, fig.115♂. (Nan: Ban Nam Poon)
Phaedyma columella martabana ; Ek-Amnuay,[2007] : 398, pl.173, figs.N90a,♂,♂(Un),♀,♀(Un). (Tham Than Lod / Krabi)
Phaedyma columella columella ; Ek-Amnuay,[2007] : 398, pl.173, figs.N90b,♂,♂(Un),♀(Malaysia [ssp.singa]).
Phaedyma columella martabana ; Ek-Amnuay,2012 : 410, pl.178, figs.N266,♂,♂(Un),♀,♀(Un). (Chiang Dao / Tham Than Lod / Krabi)
Phaedyma columella martabana ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Phaedyma columella martabana ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Ratchaburi / Saraburi) (accessed: 2020.10.05)
Phaedyma columella martabana ; Kimura et al.,2016 : 84, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao / Doi Suthep / Bun Kahan / Phu Khiewo / Muok Leck / Saraburi / Nakohn Nayok / Sarika / Wan Nam Khiewo / Kao Soi Dao / Kao Khiewo / THung Yai / Sai York Noi / Kuwai Yai / Kaeng Chai)

[ LAOS ]
Neptis columella f.alesia ; Dubois & Vitalis de Salvaza,1924 : 40. (Ban-Na-Han / Vientiane / Ban-Soui)
Phaedyma columella martabana ; Motono & Negishi,1989 : 58, pl.60, figs.1♂,2♂(Un),3♀,4♀(Un). (Vientiane / Luang Prabang / Pakse)
Phaedyma columella martabana ; Osada, Uémura & Uehara,1999 : 206, pl.56, figs.♂,♂(Un),♀, pl.57, figs.♂,♂(Un),♀. (Xaignabouri / Vientiane / Nam Tha)
Phaedyma columella martabana ; Onodera,2009b : 41. (Khammouan: Nacay)
Phaedyma columella martabana ; Nakamura & Wakahara,2012 : 45.

[ CAMBODIA ]
Neptis columella martabana ; Woodfield & Murton,2006 : fig,♂. (Botum Sakor NP.)
Phaedyma columella martabana ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)

[ VIETNAM ]
Neptis columella martabana ; Fruhstorfer,1912 : 615. (Annam)
Neptis martabana ; Dubois & Vitalis de Salvaza,1919 : 229. (Annam)
Neptis alesia ; Dubois & Vitalis de Salvaza,1919 : 229. (Annam)
Neptis columella f.martabana ; Dubois & Vitalis de Salvaza,1924 : 40. (Annam)
Neptis columella f.alesia ; Dubois & Vitalis de Salvaza,1924 : 40. (S.Annam)
Neptis columella martabana f.martabana ; Metaye,1957 : 96. (C)
Neptis columella martabana f.alesia ; Metaye,1957 : 96. (C)
Neptis columella martabana ; Metaye,1958b : 175. (Saigon, Ho Chi Minh)
Phaedyma columella martabana ; Eliot,1969 : 120. (S)
Phaedyma columella ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Huong Thuy; Phong Dien; Bach Ma)
Phaedyma columella martabana ; Miyazaki, Saito & Saito,2006b : 22, figs.N-56,♂,♀. (Lam Dong: Dambri / Dong Nai: Thac Mai)
Phaedyma columella martabana ; Monastyrskii & Devyatkin,2015 : 38. (C / S) (in part)
Phaedyma columella martabana ; Saito et al.,2017 ; 81. (S: Nui Cam)
Phaedyma columella martabana ; Monastyrskii,2019 : 4(2):87-88, pl.54, 1a,♂,1b♂(Un),1c,♂,1d,♂(Un),1e,♀,1f.♀(Un). (C: Pu Mat ; Vu Quang ; Phong Nha Ke Bang ; Phong Dien ; Huong Thuy ; Dakrong ; Bach Ma ; Kon Ka Kinh ; Yok Don ; Krong Trai ; Ninh Son ; Da Teh ; Cat Loc ; Bu Gia Map / S: Nam Cat Tien ; Lo Go Sa Mat ; La Nga)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 4♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 10♂ 6♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lamphun : 1♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 1♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Uttaradit : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Surat Thani (Koh Phangan dist.) : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Vientiane : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Champasak : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Bin Tuan : 1♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Dong Nai : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is very common in montane and secondary forest at low to moderate elevations (alt.400m - 700m).
The species is highly variable and individual and seasonal variation seems to be more pronounced than geographic variation. Further study is needed about the continental taxa.


Literature

Moore, F.,1881 : Descriptions of new Asiatic diurnal Lepidoptera.
Trans. ent. Soc. Lond. 29(3):305-313.

Fruhstorfer, H.,1905 : Neue Neptis (Phaedyma Lokalrassen.
Ent. Zeit. 19(17):89-90, 3 figs.


Phaedyma columella martabana ♂ Phaedyma columella martabana ♀
♂ (dsf), Chiang Mai, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Phaedyma columella martabana ♀+♂ Phaedyma columella martabana ♂+♀
♀+♂, Lamphun, Thailand. ♂+♀, Lampang, Thailand.
Phaedyma columella martabana ♂ Phaedyma columella martabana ♂
♂ (dsf), Chiang Mai, Thailand. ♂ (dsf), Chiang Mai, Thailand.
Phaedyma columella martabana ♂ Phaedyma columella martabana ♂
♂ (dsf), Chiang Mai, Thailand. ♂ (dsf), Chiang Mai, Thailand.

2006.02.20 - 2020.10.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.