Polyura athamas athamas  (Drury, [1773])

Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♀ Up. Polyura athamas athamas ♀ Un.
♂, Nan, Thailand. ♀, Phrae, Thailand.
Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♀ Up. Polyura athamas athamas ♀ Un.
♂, Nan, Thailand. ♀, Nan, Thailand.
Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un.
♂, Chanthaburi, Thailand. ♂, Phetchaburi, Thailand.
Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♂ genitalia Polyura athamas athamas ♂ genitalia
♂, Nan, Thailand. ♂ genitalia, Chiang Mai, Thailand.

Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un.
♂, Nakhon Si Thammarat, Thailand. ♂, Yala, Thailand.
Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un.
♂, Ranong, Thailand. ♂, Ranong, Thailand.
Polyura athamas athamas ♂ Up. Polyura athamas athamas ♂ Un. Polyura athamas athamas ♂ genitalia Polyura athamas athamas ♂ genitalia
♂, Yala, Thailand. ♂ genitalia, Ranong, Thailand.

ON : Papilio athamas Drury, [1773]
OD : Ill. nat. Hist. Exot. Insects 1: index,5, pl.2, fig.4,♂,♂(Un).
TL : China.
Distribution : Myanmar, Thailand, Laos, Cambodia, Vietnam, Tibet,
Hainan, S.China (Yunnan, Sichuan, Guangxi, Guangdong), Langkawi,
W.Malaysia.
Polyura athamas athamas map

Synonyms

Charaxes samatha Moore, [1879] : Proc. zool. Soc. Lond. 1878(4):831. TL. Moolai, Tenasserim.
Eriboea athamas dexippus Fruhstorfer, 1914 : Ent. Rundschau 31(1):2. TL. Tai Ninh. (Geneva Mus.)


Records taken from Literature

[ THAILAND ]
Eriboea athamas samatha ; Fruhstorfer,1914 : 719. (Ranong; Petchaburi)
Eulepis athamas ; Godfrey,1916 : 133.
Eriboea athamas samatha ; Pendlebury,1923 : 42. (Nakhon Sri Thammarat: Lan Saka; Khao Luang)
Eriboea athamas samatha ; Godfrey,1927 : 124. (N)
Eriboea athamas samatha ; Godfrey,1930 : 301. (except P)
Eriboea athamas samatha ; Godfrey,1932a : 270. (P)
Polyura athamas samatha ; Pinratana,1979 : 98, figs.N166, ♂,♂(Un). (Chanthaburi; Chonburi; Kanchanaburi; Nakhon Nayok; Nonthaburi; Ratchaburi; Trang)
Polyura athamas athamas ; Smiles,1984 : 167. (Doi Inthanon; [Klong Pong Kapo]; [Hue Tak So]; [Muok-Lek]; Phrae: [Me Sai Song]; Khlong Khlung; [Khao Sabab]; Chanthaburi; Hin Lap; Bangkok; Ranong; Prauchuap; [Pak Tawan])
Polyura athamas athamas ; Pinratana & Eliot,1996 : 98, pl.75, figs.187,♂,♀. (Chiang Mai; Chaiyaphum; Nakhon Ratchasima; Nakhon Nayok; Chonburi; Chanthaburi; Kanchanaburi; Ratchaburi; Trang; Yala)
Polyura athamas athamas ; Yamaguchi & Aoki,2000 : 256, fig.115,♂. (Mukdahan: Phu Sa Duak Bua; Phu Mu)
Polyura athamas athamas ; Yamaguchi, Aoki & Akiyama,2005 : 52. (Ubon Ratchathani: Phu Pung)
Polyura athamas athamas ; Katsuyama et al.,2005 : 71, fig.177,♂. (Umphang)
Polyura athamas athamas ; Katsuyama et al.,2006 : 50, fig.125,♂. (Nan: Ban Nam Poon)
Polyura athamas athamas ; Ek-Amnuay,[2007] : 486, pl.217, figs.N200,♂,♂(Un),♂,♂(Un),♀[♂],♀(Un)[♂(Un)]. (Kanchanaburi; Rangsit)
Polyura athamas athamas ; Les Day,2011- : Samui Butterflies, figs. (Koh Samui) (accessed: 2021.07.01)
Polyura athamas athamas ; Ek-Amnuay,2012 : 358, pl.152, figs.N188,♂,♂(Un),♂,♂(Un),♀,♀(Un). (Rangsit; Kanchanaburi)
Polyura athamas athamas ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Polyura athamas athamas ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, fig. (Nakhon Ratchasima) (accessed: 2021.07.01)
Polyura athamas athamas ; Kimura et al.,2016 : 132-133, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un). (Chiang Dao; Doi Inthanon; Doi Pa Chang; Thung Saleng Luang; Phu Khiewo; Wan Nam Khiewo; Kao Soi Dao; Kratchin WF; Pliewn WF; Kao Khiewo; Muang Shone; Khlong Naka; Than To WF.)

[ LAOS ]
Eriboea athamas athamas ; Fruhstorfer,1914 : 718. (Luang Prabang)
Eriboea athamas ; Dubois & Vitalis de Salvaza,1919 : 233. (Laos)
Polyura athamas athamas ; Smiles,1984 : 167. ([Luhang Prahang]; [Muang Baw]; Sekong River)
Polyura athamas samatha ; Motono & Negishi,1989 : 62, pl.77, figs.3,♂,4,♂(Un). (Vang Vieng; Phatong; Tha Ngone; Vientiane; Luang Prabang; Ban Houei Sai; Nam Ngum Dam)
Polyura athamas athamas ; Osada, Uémura & Uehara,1999 : 208, pl.78, figs.♂,♂(f.hamasta),♂(Un). (Vang Vieng; Oudomxay; Nong Het)
Polyura athamas athamas ; Nimura & Wakahara,2005 : 44. (Namdhua; Thakhek; Nhahin)
Polyura athamas athamas ; Onodera,2009b : 35,36,39,43. (Vientiane: Namsong Riverside; Xieng Kouang: Muang Kham; Bolikhamxai: Thabok; Phon Si)
Polyura athamas athamas ; Nakamura & Wakahara,2012 : 47.

[ CAMBODIA ]
Polyura athamas athamas ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)

[ VIETNAM ]
Eriboea athamas athamas ; Fruhstorfer,1914 : 718. (Tonkin / S.Annam)
Eriboea athamas ; Dubois & Vitalis de Salvaza,1919 : 233. (Annam / Tonkin)
Eriboea athamas athamas f.aest.athamas ; Metaye,1957 : 99. (N / C / S)
Eriboea athamas athamas f.hib.hamasta ; Metaye,1957 : 99. (N / C / S)
Polyura athamas athamas ; Smiles,1984 : 167. (Muong Khuoug; Tongkin: Yen Bai; Chiem Hoa; [Nam-Hou]; Xom Giong; [Nacham]; [Bac-Kan])
Polyura athamas athamas ; Ikeda et al.,2000 : 37, figs.8:5,♂,6,♂(Un), fig.9:46(♂,genitalia). (Cuc Phuong)
Polyura athamas athamas ; Luong et al.,2004 : 8, 71, figs.♂,♂(Un). (Cuc Phuong)
Polyura athamas athamas ; Miyazaki, Saito & Saito,2006b : 34, fig.N-111,♂. (Lam Dong: Dai Duc Me; Madagui; Dong Nai: Thac Mai)
Polyura athamas ; Monastyrskii et al.,2006 : 42. (Thua Thien Hue: Nam Dong; A Luoi; Phong Dien; Bach Ma)
Polyura athamas athamas ; Miyazaki et al.,2010 : 24, fig.N-111,♀. (Lam Dong: Dambri)
Polyura athamas athamas ; Monastyrskii & Devyatkin,2015 : 42. (N / C / S)
Polyura athamas athamas ; Monastyrskii,2019 : 4(1):85-86, pl.12, figs.1a♂,1b♂(Un),1c♀. (N: Hoang Lien; Ba Be; Tam Dao; Cuc Phuong; Pu Luong / C: Xuan Lien; Ben En; Pu Hoat; Bu Huong; Pu Mat; Huong Son; Vu Quang; Phong Dien; Dakrong; Bach Ma; Nam Dong; A Luoi; Que Phuoc; Kon Ka Kinh; Chu Yang Sin; Yok Don; Bu Gia Map; Nui Chu / S: Cat Tien; Lo Go Sa Mat; La Nga)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 12♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 6♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 1♂ (photo by S.Sophonviwatkul).
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Ranong : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Si Thammarat : 3♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Yala : 5♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Xaignabouli : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is common in montane forest at low to moderate elevations (alt.30m - 1300m). Males are territorial on shrubs or trees.
Toussaint,2015 described Polyura paulettae by mainly DNA analysis. I suppose it may be widely distributed in Indo-China including P.Thailand. It is highly probable that this species is present in records of athamas in this area. However I treat the records in the previous literatures as athamas for the time being.
Male genitalia has small difference between examples from continental area and Peninsular Thailand. Further study is needed to the relationship of these.


Literature

Drury, D.,[1773] : Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1.
1-130, pl. 1-50.

Moore, F.,[1879] : A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim,
with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4):821-859, 3 pls.

Fruhstorfer, H.,1914 : Rhopaloceren von Cochinchina
Ent. Rundschau 31(1):1-3.

Smiles, R.L.,1982 : The taxonomy and phylogeny of the genus Polyura Billberg (Lepidoptera: Nymphalidae)
Bull. Brit. Mus. Ent. (Nat. Hist.) 44(3):115-237. London.


Polyura athamas athamas ♂ Polyura athamas athamas ♂
♂, Loei, Thailand. ♂, Chiang Mai, Thailand.
Polyura athamas athamas ♂ Polyura athamas athamas ♂
♂, Yala, Thailand. ♂, Nakhon Si Thammarat, Thailand.

2006.12.09 - 2021.07.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.