Polyura arja arja
(C. Felder & R. Felder, 1867)
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♂, Nan, Thailand. |
♀, Nan, Thailand. |
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♂, Nan, Thailand. |
♂, Chonburi, Thailand. |
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♂, Phetchaburi, Thailand. |
♂ genitalia, Nan, Thailand. |
| arja ? |
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♂, Chiang Mai, Thailand. |
♂, Vientiane, Laos. |
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♂, Chonburi, Thailand. |
♂ genitalia, Vientiane, Laos. |
| f. roeberi |
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♂, Tak, Thailand. |
♀, Kachin, Myanmar. |
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♂, Kachin, Myanmar. |
♂ genitalia, Tak, Thailand. |
ON :
Charaxes arja C. Felder & R. Felder, 1867
OD : Reise öst. Fregatte Novara. Zool. Theil (3):438.
TL : 'India septentrionalis: Assam' [Sylhet, India]. (NHML)
Distribution : Sikkim, Assam to Myanmar, Thailand, Laos, Cambodia, Vietnam, Yunnan.
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Synonyms
Charaxes arja roeberi Fruhstorfer, 1898 : Ent. Nachr. 24(4):59. TL. "Khasia Hills" [Khasi Hills, Meghalaya, India].
Eulepis arja f. temp. vernus Rothschild, 1899 : Novit. zool. 6:244-245. TL. Sikkim. (NHML) (IFS)
Type Material Information
arja :
Lectotype ♂, India, Silhet. designated by R.L.Smiles,1979. (NHML)
Paralectotype 1♂, India Assam. designated by R.L.Smiles,1979.(NHML)
vernus : Holotype ♂, 23.Mar.1888. Sikkim. (NHML)
Records taken from Literature
[ THAILAND ]
Eriboea arja ; Fruhstorfer,1914 : 720. (Petchaburi)
Eulepis arja ; Godfrey,1916 : 133. (Chonburi: Sriracha)
Eriboea arja ; Godfrey,1927 : 124. (N)
Eriboea arja ; Godfrey,1930 : 301. (Prae: Me Song forest; Chiang Rai; Muak Lek; Chonburi: Sriracha; Me Wong)
Polyura arja arja ; Pinratana,1979 : 98, figs.N167,♂,♂(Un). (Chonburi; Nan)
Polyura arja arja ; Smiles,1984 : 177. (Pak Jong; Mae Wong; [Me Tharn]; Mae Sariang; Hin Lap; [Muok-Lek]; [Pang Yao]; [Raheng])
Polyura arja arja ; Pinratana & Eliot,1996 : 98, pl.75, figs.188,♂,♀. (Chiang Mai; Chaiyaphum; Phisanulok; Nakhon Ratchasima; Nakhon Nayok; Chonburi; Chanthaburi; Saraburi; Kanchanaburi)
Polyura arja arja ; Yamaguchi & Aoki,2000 : 256, fig.116,♂. (Mukdahan: Phu Pha Thaep; Phu Mu)
Polyura arja arja ; Yamaguchi & Aoki,2002 : 31. (Mukdahan: Ban Lao Kram; Ban Na Lak; Ubon Ratchathani: Kaeng E-Keow)
Polyura arja arja ; Yamaguchi, Aoki & Akiyama,2005 : 52. (Ubon Ratchathani: Phu Pung; Mukdahan: Ban Lao Kram)
Polyura arja arja ; Katsuyama et al.,2005 : 71, fig.178,♂. (Umphang: Tha-Sai; Umphang)
Polyura arja arja ; Katsuyama et al.,2006 : 50, fig.124,♂. (Nan: Ban Nam Poon)
Polyura arja arja ; Ek-Amnuay,[2007] : 486, pl.217, figs.N201,♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao)
Polyura arja arja ; Ek-Amnuay,2012 : 358, pl.152, figs.N189,♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao)
Polyura arja arja ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs, Early Stages. (Chiang Mai; Chanthaburi) (accessed: 2023.05.04)
Polyura arja arja ; Kimura et al.,2016 : 133, figs.♂,♂(Un),♀,♀(Un). (Chiang Mai; Maethro; Thung Saleng Luang; Phu Khiewo; Nakohn Nayok; Saraburi; Wan Nam Khiewo; Kao Soi Dao; Pliewn WF; Kao Khiewo; Sai York Noi)
[ LAOS ]
Eriboea arja f. vernus ; Dubois & Vitalis,1924 : 46. (Ban-Samoui)
Eriboea arja f. roeberi ; Dubois & Vitalis,1924 : 46. (Nam-Tie)
Polyura arja arja ; Motono & Negishi,1989 : 62, pl.77, figs.5,♀,6,♀(Un). (Vang Vieng; Phatong; Nam Moh; Tha Ngone; Vientiane; Luang Prabang; Ban Houei Sai)
Polyura arja arja ; Osada, Uémura & Uehara,1999 : 208, pl.78, figs.♂,♂(f. roeberi),♀,♀,♂(Un). (Oudomxay; Vientiane; Nong Het)
Polyura arja arja ; Nimura & Wakahara,2005 : 43-44. (Namdhua; Thakhek)
Polyura arja arja ; Onodera,2009b : 39. (Bolikhamxai: Thabok)
Polyura arja arja ; Nakamura & Wakahara,2012 : 47.
Polyura arja arja ; Onodera,2022 : 50, pl.148, figs.♂,♂(Un). (Khammouan: Nahin)
[ CAMBODIA ]
Polyura arja arja ; Woodfield & Murton,2006 : fig,♂. (Botum Sakor NP.)
Polyura arja arja ; Kosterin,2019b : 32, figs.30,♂(Un),31,♂(Un). (Ratanakiri; Mondulkiri)
Polyura arja arja ; Chartier & Kosterin,2022 : 117. (Koh Kong; Mondulkiri; Ratanakiri; Siem Reap)
[ VIETNAM ]
Eriboea arja ; Fruhstorfer,1914 : 720. (Tonkin: Chiem Hoa)
Eriboea arja ; Vitalis,1919 : 233. (Tonkin)
Eriboea arja f. arya [sic] ; Dubois & Vitalis,1924 : 46. (Tonkin: Chiem-Hoa)
Eriboea arja roeberi ; Metaye,1958b : 174. (Dong Nai: Trang Bom)
Polyura arja arja ; Smiles,1984 : 177. (Chiem Hoa; [Nam-Hou]; [Ko-Tich]; Van Bu; Cha Pa)
Polyura arja arja ; Ikeda et al.,2000 : 37, figs.8:7,♂,8,♂(Un), fig.9:47(♂,genitalia). (Cuc Phuong)
Polyura arja ; Luong et al.,2004 : 8. (Cuc Phuong)
Polyura arja arja ; Miyazaki, Saito & Saito,2006b : 34, fig.N-112,♂. (Lam Dong: Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Polyura arja arja ; Miyazaki, Saito & Saito,2007c : 38, fig.N-112,♀. (Lam Dong: Dambri)
Polyura arja arja ; Nihira,2009 : 99, fig.♂. (Phu Quoc)
Polyura arja arja ; Monastyrskii & Devyatkin,2015 : 42. (N / C / S)
Polyura arja arja ; Monastyrskii,2019 : 4(1):87-88, pl.13, figs.2a♂,2b♂(Un),2c♀. (N: Muong Nhe; Huu Lien ; Cuc Phuong / C: Pu Mat; Nui Chua; Yok Don; Chu Yang Sin; Cat Tien / S: Cat Tien; Binh Chau)
Specimens Examined
[ THAILAND ]
Mae Hong Son : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Chiang Mai : 10♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Nan : 2♂ 4♀.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Chonburi : 2♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Phetchaburi : 2♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
[ VIETNAM ]
Lam Dong : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Dong Nai : 1♀.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
< arja ? >
[ THAILAND ]
Chiang Mai : 2♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Chonburi : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
[ LAOS ]
Vientiane : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
< f. roeberi >
[ THAILAND ]
Tak : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
[ MYANMAR ]
Kachin (N.E.Putao) : 3♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Notes
The butterfly is common in montane forests at low to moderate elevations (Alt. 300 - 900m).
I suspect Polyura arja has several cryptic species.
Further taxonomic study is needed.
Literature
Felder, C. & Felder, R.,[1865]-1874 : Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair.
Zoologischer Theil. Zweiter Band. Zweite Abtheilung: Lepidoptera. Wien, Carl Gerold's Sohn.
(1):[4] + 1-136, pls. 1-21 ([12 January 1865]) ; (2):[2] + 137-378, pls. 22-47 ([after October] 1865) ; (3):[2] + 379-536, pls.48-74 ([25 April] 1867) ; (4):[6] + 1-9 [explanation of plates] ([31 December] 1874)
Fruhstorfer, H.,1898 : Ueber einige Charaxes-Arten.
Ent. Nachr. 24(4):53-60. ([15].Feb.1898)
Rothschild, L.W. & Jordan, K.,1899 : A monograph of Charaxes and the allied Prionopterous genera. Part 2.
Novit. zool. 6:220-286, pl.7.
Smiles, R.L.,1982 : The taxonomy and phylogeny of the genus Polyura Billberg (Lepidoptera: Nymphalidae)
Bull. Brit. Mus. Ent. (Nat. Hist.) 44(3):115-237. London.
Lang, S.Y.,2009 : A Study on the systematics of Nymphalidae (excepting Satyrinae) from China (Lepidoptera: Papilionoidea)
Beijing, P.R.China.
Lang, S.Y.,2012 : The Nymphalidae of China (Lepidoptera, Rhopalocera) Part 1
Libytheinae, Danainae, Calinaginae, Morphinae, Heliconiinae, Nymphalinae, Charaxinae, Apaturinae, Cyrestinae, Biblidinae, Limenitinae
456 pp, 32 col. + 41 black&white pls. Tshikolovets Publications, Pardubice.
Fang, L. & Zhang, Y.,2018 : Review of the genus Polyura Billberg (Lepidoptera: Nymphalidae) from China with description of a new species
Entomotaxonomia 40(1):27-45.
A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.
