Leptosia nina nina  (Fabricius, 1793)

Leptosia nina nina ♂ Up. Leptosia nina nina ♂ Un. Leptosia nina nina ♀ Up. Leptosia nina nina ♀ Un.
♂, Mae Hong Son, Thailand. ♀, Mae Hong Son, Thailand.
Leptosia nina nina ♂ Up. Leptosia nina nina ♂ Un. Leptosia nina nina ♀ Up. Leptosia nina nina ♀ Un.
♂, Bangkok, Thailand. ♀, Ranong, Thailand.
Leptosia nina nina ♂ Up. Leptosia nina nina ♂ Un. Leptosia nina nina ♀ Up. Leptosia nina nina ♀ Un.
♂, Yala (Than To dist.), Thailand. ♀, Yala (Than To dist.), Thailand.
Leptosia nina nina ♂ Up. Leptosia nina nina ♂ Un. Leptosia nina nina ♂ Up. Leptosia nina nina ♂ Un.
♂, Chanthaburi, Thailand. ♂, Satun, Thailand.

ON : Papilio nina Fabricius, 1793
OD : Ent. syst. 3(1):194-195.
TL : India orientali.
Distribution : Sri Lanka, India to Myanmar,
Thailand, Laos, Vietnam, Hainan, S.China,
Langkawi, W.Malaysia (Kedah; Penang).
● ssp.nina
transition zone of nina and malayana
ssp.malayana
Leptosia nina nina map

Synonym

Papilio xiphia Fabricius, 1781 : Spec. Ins. 2:43. TL. India orientali. (homonym)


Records taken from Literature

[ THAILAND ]
Leptosia xiphia xiphia ; Fruhstorfer,1910 : 121. (Siam)
Leptosia xiphia xiphia ; Godfrey,1916 : 113.
Leptosia xiphia xiphia ; Pendlebury,1923 : 25. (Nakhon Sri Thammarat: Tai Sai River)
Leptosia xiphia xiphia ; Godfrey,1927 : 120. (N)
Leptosia xiphia xiphia ; Godfrey,1930 : 220-221. (all over)
Leptosia xiphia xiphia ; Godfrey,1931 : 192. (Ko Tao)
Leptosia xiphia xiphia ; Godfrey,1932a : 264. (P)
Leptosia nina nina ; Pinratana,1983 : 10, pl.9, figs.15a♂,♂(Un). (Chanthaburi; Chaiyaphum; Kanchanaburi; Nakhon Nayok; Lop Buri; Nakhon Ratchasima; Nonthaburi; Petchaburi; Saraburi; Phang Naga; Nakhon Sri Thammarat; Songkhla)
Leptosia nina nina ; Yamaguchi & Aoki,2000 : 251-252, fig.46,♂. (Mukdahan: Phu Mu)
Leptosia nina nina ; Yamaguchi & Aoki,2002 : 18. (Mukdahan: Ban Lao Kram; Ban Gok Tum)
Leptosia nina nina ; Yamaguchi, Aoki & Akiyama,2005 : 46. (Mukdahan: Nam Tok Tad Ton; Ubon Ratchathani: Don Far Huan)
Leptosia nina nina ; Katsuyama et al.,2005 : 65, fig.48,♂. (Umphang: Tha Sai; Umphang)
Leptosia nina nina ; Ek-Amnuay,[2007] : 138, pl.41, figs.Pi15a♂,♂(Un),♀,♀(Un). (Bangkok; Chiang Mai)
Leptosia nina malayana ; Ek-Amnuay,[2007] : 138, pl.41, figs.Pi15b,♂[Un],♂(Un)[Up]. (Yala; Khao Phanom Bencha)
Leptosia nina nina ; Les Day,2011-2022 : Samui Butterflies, figs. (Koh Samui) (accessed: 2022.04.16)
Leptosia nina nina ; Kimura et al.,2011 : 154, figs.♂,♂(Un),♀,♀(Un). (Sarika; Nakohn Nayok; Wan Nam Khiewo; Phu Khiewo; Kao Soi Dao; Kanchanaburi; Khlong Naka; Muang Shone; Haadjai; Satun)
Leptosia nina malayana ; Kimura et al.,2011 : 154, figs.♂,♂(Un),♀,♀(Un). (Than To WF.)
Leptosia nina nina ; Ek-Amnuay,2012 : 138, pl.41, figs.Pi15a♂,♂(Un),♀,♀(Un). (Bangkok; Chiang Mai)
Leptosia nina nina ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Bangkok; Saraburi; Chanthaburi) (accessed: 2022.04.16)
Leptosia nina malayana ; Küppers & Janikorn,2017 : 191, figs.Abb.Pi-024,033-034,118-122. (Phang Nga)
Leptosia nina nina ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Leptosia nina nina ; Motono & Negishi,1989 : 48, pl.15, figs.1,♂,2,♂(Un). (Tha Ngone; Vientiane)
Leptosia nina nina ; Osada, Uémura & Uehara,1999 : 202, pl.31, figs.♂,♀. (Thakhek; Nhahin)
Leptosia nina nina ; Nimura & Wakahara,2005 : 39. (Namdhua; Thakhek)
Leptosia nina nina ; Onodera,2009b : 34-35. (Vientiane: Namsong Riverside)
Leptosia nina nina ; Nakamura & Wakahara,2012 : 42.

[ CAMBODIA ]
Leptosia nina nina ; Woodfield & Murton,2006 : figs,♂,♂(Un). (Botum Sakor NP.)
Leptosia nina ; Onodera,2007 : 21,29, fig.10,♂. (Rattanakiri: Ta Veaeng; Siem Reap: Angkor)
Leptosia nina ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)

[ VIETNAM ]
Leptosia xiphia xiphia ; Fruhstorfer,1910 : 121. (Annam)
Leptosia xiphia xiphia ; Godfrey,1919 : 464. (S: Daban)
Leptosia xiphia xiphia ; Moulton,1923 : 132. (Pulo Condore)
Leptosia xiphia ; Lemee,1950 : 63. (Saigon)
Leptosia xiphia xiphia ; Metaye,1957 : 87. (N / C / S)
Leptosia nina nina ; Ikeda et al.,1998 : 19, figs.6:19,♂,10:33(♂genitalia). (Cuc Phuong)
Leptosia nina ; Vu,2003 : 56. (Phu Quoc)
Leptosia nina ; Luong et al.,2004 : 2, 30, figs.♂,♂(Un). (Cuc Phuong)
Leptosia nina ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Leptosia nina nina ; Miyazaki, Saito & Saito,2006 : 11, figs.Pi-6,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien; Ta Cu; Ho Chi Minh)
Leptosia nina nina ; Nihira,2009 : 98. (Phu Quoc)
Leptosia nina nina ; Monastyrskii & Devyatkin,2015 : 16. (N / C / S)
Leptosia nina nina ; Saito et al.,2017 ; 78. (S: Nui Cam; Nui Coto; Nui Ket)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Rai : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 12♂ (incl.photo & photo by M.Sugiyama).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 5♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Kanchanaburi : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Nayok : 2♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Sa Kaeo : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Bangkok : 3♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Trat : 1♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Ranong : 9♂ 3♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Surat Thani (Koh Phangan Dist.) : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Krabi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Satun (Koh Tarutao) : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Yala (Than To Dist.) : 20♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Pakse : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is very common everywhere at low to high elevations (Alt. 10 - 1800m).
The specimens from Than To, Yala have been considered as belonging to ssp.malayana, however after comparing a large number of specimens, they should be included in the nominotypical subspecies. The population is generally ssp.nina, however, individuals similar to ssp.malayana also occur rarely. It seems to be a transition zone of both subspecies.


Literature

Fabricius, J.C.,1781 : Species Insectorum exhibentes eorum differentias specificas, synonyma auctorum, locanatalia, metamorphosin adjectis observationibus descriptionibus.
vol.2:1-494. Humburg and Cologne, Karl E.Bohn.

Fabricius, J.C.,1793 : Entomologia systematica emedata et aucta. secundum classes, ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus.
3(1): [iv] + 488 pp. Copenhagen. C.G.Proft.


Leptosia nina nina ♂ Leptosia nina nina ♂
♂, Chiang Mai, Thailand. (photo by M.Sugiyama) ♂, Chiang Mai, Thailand.
Leptosia nina nina ♂ Leptosia nina nina ♂
♂+♀ (mating), Trat, Thailand.

1996.10.31 - 2022.05.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.