Graphium arycles arycles  (Boisduval, 1836)

Graphium arycles arycles ♂ Up. Graphium arycles arycles ♂ Un. Graphium arycles arycles ♀ Up. Graphium arycles arycles ♀ Un.
♂, Chanthaburi, Thailand. ♀, Trat, Thailand.
Graphium arycles arycles ♂ Up. Graphium arycles arycles ♂ Un. Graphium arycles arycles ♀ Up. Graphium arycles arycles ♀ Un.
♂, Vientiane, Laos. ♀, Houaphan, Laos. (photo by K.Saito)
Graphium arycles arycles ♀ Up. Graphium arycles arycles ♀ Un. Graphium arycles arycles ♂ genitalia Graphium arycles arycles ♂ genitalia
♀, Bolikhamxai, Laos. (photo by A.M.Cotton) ♂ genitalia, Bolikhamxai, Laos.

ON : Papilio arycles Boisduval, 1836
OD : Spec. gen. Lepid. 1:231-232.
TL : 'Java or Sumatra' ['Annam, Cochinchina or Camboja']
Distribution : S.E & E.Thailand, Laos, Cambodia, Vietnam.
[? Manipur ; Myanmar]
● ssp.arycles
ssp.rama
Graphium arycles arycles Map

Synonyms

Papilio arycles sphinx Fruhstorfer, 1899 : Soc. ent. 14(7):49. TL. ? loc. [Indochina]. (♂, NHML)
Papilio arycles arycleoides Fruhstorfer, [1902] : Deut. ent. Zeit. [Iris] 14(2):344. TL. Muok-Lek, Siam. (♂, NHML)
Graphium arycles masal Kocak & Kemal, 2000 : Cent. ent. Stud. Misc. 71:5. (Unneccessary replacement name for Papilio arycles sphinx)
Arisbe arycles occidentalis Page & Treadaway, 2014 : Stutt. Beit. Natur. Serie A, Neue Ser. 7:277-278, figs.27(HT,♂),28(HT,♂Un). TL. 'Chiang Mai', Thailand.


Type Materials Information

arycles ; '♂, Type (red), #149417.' (NHML, examined) [This is not a true type specimen.]
sphinx ; ♂, Paratype (yellow), Indochina. H.Fruhstorfer (NHML, examined)
'♂, Type HT (red), #149418, Indochina. H.Fruhstorfer' (NHML, examined) ; '♀ Type, Tay-Ninh, Cochinchina. Fruhstorfer' (NHML, examined) [These are not type specimens.]
arycleoides ; ♂, Type HT (red), #149416, Muok-Lek, Siam. Januar. Fruhstorfer (NHML, examined)
occidentalis ;
Holotype ♂, May.1966. Chiang Mai, Thailand (SMF).
Paratypes : 2♂, May.1966. Chiang Mai, Thailand. (CGT) ; 1♂, S.Myanmar (MGP) ; 1♂, Manipur, N.E.India. (MGP)


Records taken from Literature

[ THAILAND ]
Papilio arycles arycleoides ; Fruhstorfer,[1903] : 220-221. (Muok-Lek)
Papilio arycles arycleoides ; Jordan,1909 : 102. (Muok Lek)
Papilio arycles arycleoides ; Godfrey,1916 : 112. (Nong Khor)
Papilio arycles arycleoides ; Godfrey,1930 : 218-219. (Muak Lek; Chonburi: Sriracha)
Graphium arycles arycleoides ; Pinratana,1977 : 59, figs.P49a♀,♂(Un). (Wangtrakrai; Chonburi: Sriracha)
Graphium arycles arycleoides ; Pinratana & Eliot,1992 : 37,46, pl.68, figs.61b♂,♂(Un). (Chanthaburi; Chonburi; Nakhon Ratchasima)
Graphium arycles arycleoides ; Yamaguchi & Aoki,2000 : 251, fig.43,♂. (Mukdahan: Phu Mu)
Graphium arycles arycleoides ; Yamaguchi & Aoki,2002 : 16. (Mukdahan: Ban Lao Kram; Ubon Ratchathani: Nam Tok Soi Sawan; Kaeng E-Keow; Phu Hin Dang)
Graphium arycles arycleoides ; Yamaguchi, Aoki & Akiyama,2005 : 46. (Ubon Ratchathani: Phu Chong Na Yoi; Mukdahan: Ban Lao Kram; Phu Hin Khan)
Graphium arycles sphinx ; Ek-Amnuay,[2007] : 122, pl.33, figs.P70b♂,♂(Un),♂(Un). (Chanthaburi)
Graphium arycles arycleoides ; Kimura et al.,2011 : 139, figs.♂,♂(Un),♀,♀(Un). (Saraburi; Wan Nam Khiewo; Khao Khiewo; Khlong Tariew)
Graphium arycles sphinx ; Ek-Amnuay,[2007] : 120, pl.33, figs.P67b♂,♂(Un),♂(Un). (Chanthaburi; Pang Sida)
Arisbe arycles arycles ; Page & Treadaway,2014 : 276. (E)
Arisbe arycles arycleoides ; Page & Treadaway,2014 : 277. (Chanthaburi)
Arisbe arycles occidentalis ; Page & Treadaway,2014 : 277-228, figs.27(HT,♂),28(HT,♂Un). ('Chiang Mai')
Graphium arycles arycles ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Sa Kaeo; Saraburi; Chanthaburi) (accessed: 2022.10.20)

[ LAOS ]
Papilio arycles f.arycles ; Dubois & Vitalis,1921 : 16. (Laos)
Graphium arycles arycleoides ; Motono & Negishi,1989 : 47, pl.10, figs.5,♂,6,♂(Un). (Vientiane; Tha Ngone; Nam Ngum Dam)
Graphium arycles arycleoides ; Osada, Uémura & Uehara,1999 : 201, pl.27, figs.♂,♂(Un),♀. (Thabok; Phong Si)
Graphium arycles sphinx ; Cotton & Racheli,[2007] : 309-310. (Thabok; Hinboun; Lak Sao)
Graphium arycles sphinx ; Onodera,2009b : 39-40. (Bolikhamxai: Thabok; Khammouan: Thakhek)
Graphium arycles sphinx ; Nakamura & Wakahara,2012 : 42.
Arisbe arycles arycles ; Page & Treadaway,2014 : 276. (Laos)

[ CAMBODIA ]
Graphium arycles ; Onodera,2008 : 59, figs.♂,♂(Un).
Graphium arycles ; Onodera,2009 : 37. (Kratie: Snoul)
Arisbe arycles arycles ; Page & Treadaway,2014 : 276. (Cambodia)

[ VIETNAM ]
Papilio arycles sphinx ; Fruhstorfer,[1903] : 220-221. (Tonkin ?)
Papilio arycles ; Dubois & Vitalis,1914 : 151. (Annam: Hue)
Papilio arycles f.arycles ; Dubois & Vitalis,1921 : 16. (Annam: Hue)
Papilio arycles arycles ; Metaye,1957 : 84. (C)
Graphium arycles ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; Huong Thuy; Phong Dien; Bach Ma)
Graphium arycles arycleoides ; Monastyrskii,2007 : 84-85, pl.26, fig.4,♂. (C: Bach Ma; Phong Dien; Nam Dong; Huong Thuy / S: Lo Go Sa Mat; Saigon)
Graphium arycles arycles ; Nihira,2009 : 97, fig.♂. (Phu Quoc)
Graphium arycles sphinx ; Miyazaki et al.,2010 : 20, figs.Pa-44,♂,♀. (Dong Nai: Thac Mai)
Arisbe arycles arycles ; Page & Treadaway,2014 : 276. (S)
Graphium arycles sphinx ; Monastyrskii & Devyatkin,2015 : 14. (C / S)


Specimens Examined

[ THAILAND ]
Mukdahan : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Si Saket : 1♂ (photo by A.Giudici).
JanFebMarAprMayJun JulAugSepOctNovDec

Sa Kaeo : 2♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Nayok : 2♂ (photo by T.Aoyama).
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 6♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Trat : 1♂ 1♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Vientiane : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 50♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 15♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Houaphan : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is not rare in montane forest at low elevations (alt.10m - 350m). Males frequently come to the banks of streams and puddles to suck water.
Page & Treadaway,2014 considered the type locality of arycles as Annam, Cochinchina or Cambodia. I agree with their treatment, considering the pattern of red markings on the UnH in the original description as the decisive basis.
Page & Treadaway,2014 separated continental populations to 3 subspecies such as arycles, arycleoides and occidentalis. However these are considered variations and cannot be clearly separated. The type locality of ssp.occidentalis is 'Chiang Mai'. But, as far as we know, it has never been found in Chiang Mai province or anywhere in N.W.Thailand until now. Furthermore, according to Treadaway, these specimens were bought from an insect dealer in Chiang Mai and were not collected by himself. It is most likely to be an erroneous locality, possibly from S.E.Thailand (Chanthaburi etc.).
Kimura assigned 'Khlong Tariew' to Kanchanaburi, W.Thailand, however this location name is in Chanthaburi, S.E.Thailand and there is no place in Kanchanaburi with this name. After reviewing various circumstantial evidence, including other species recorded at the same locality, we consider that this is definitely Chanthaburi in S.E.Thailand, not Kanchanaburi in W.Thailand. Further distributional research is needed.
The taxon sphinx is based on a single aberrant form in which the UnH red spots are replaced by yellow. There are several "type specimens" in the NHML. The male type (HT) and female type have red spots and are cannot be type specimens. The male paratype (yellow) has yellow spots as described in the original description. Thus this specimen may be the true type specimen.

The difference between ssp.arycles and ssp.rama.
Graphium arycles arycles UnH Graphium arycles rama UnH
arycles UnH, Trat, Thailand. rama UnH, Yala, Thailand.


Literature

Boisduval, J.B.A.,1836 : In Roret's "Suite a Buffon", Histoire naturelle des Insectes. Species general de Lepidopteres. 1.
Paris, Libr. Encycl. de Roret 8vo:[4]+xii+690+6pp, 24 pls.

Fruhstorfer, H.,1899 : Drei neue Papilios.
Soc. ent. 14(7):49.

Fruhstorfer, H.,[1902] : Neue Indo-Australische Lepidopteren.
Deut. ent. Zeit. [Iris] 14(2):334-350.

Jordan, K.,1909 : Die Indo-Australische Tagfalter. Papilionidae in Seitz, Grossschmett. Erde
9:97-104.

Kocak, A.O. & Kemal, M.,2000: Sergi Asiye ve Pasipiktiki bezi taksonlar heqqide nomenklatur hatiriliri (Papilionidae, Lepidoptera).
Cent. ent. Stud. Misc. 71:1-6, 11.

Page, M.G.P. & Treadaway, C.G.,2014 : Revisional notes on the Arisbe eurypylus species group (Lepidoptera: Papilionidea: Papilionidae)
Stutt. Beit. Natur. Serie A, Neue Ser. 7:253-284.


Graphium arycles arycles ♂ Graphium arycles arycles ♂
♂, Sa Kaeo, Thailand. ♂, Sa Kaeo, Thailand.
Graphium arycles arycles ♂ Graphium arycles arycles ♂
♂, Nakhon Nayok, Thailand. (photo by T.Aoyama) ♂ (with doson, eurypylus), Nakhon Nayok, Thailand. (photo by T.Aoyama)

1996.10.08 - 2022.10.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.