Papilio helenus helenus  Linnaeus, 1758

Papilio helenus helenus ♂ Up. Papilio helenus helenus ♂ Un. Papilio helenus helenus ♀ Up. Papilio helenus helenus ♀ Un.
♂ (ds), Tak, Thailand. ♀ (ds), Chiang Mai, Thailand.
Papilio helenus helenus ♂ Up. Papilio helenus helenus ♂ Un. Papilio helenus helenus ♀ Up. Papilio helenus helenus ♀ Un.
♂ (ws), Xaignabouri, Laos. ♀ (ws), Nan, Thailand.

ON : Papilio helenus  Linnaeus, 1758
OD : Syst.nat. (ed.10) 1:459.
TL : 'Asia'.
Distribution : Nepal, India to Myanamr, Thailand,
Laos, Cambodia, Vietnam, Hainan, S.China, Langkawi,
W.Malaysia, Tioman, Singapore.
Papilio helenus helenus Map

Synonyms

Papilio helenus ♂ab.rufatus Rothschild, 1895 : Novit. zool. 2(3):286. TL. India ; Burma ; Malacca ; China.
Papilio helenus aulus Fruhstorfer, 1908 : Ent. Wochenbl. 25:38. TL. Hainan. (NHML)
Papilio helenus ab.aurea Boullet & Le Cerf, 1912 : Bull. Soc. ent. Fr. 1912(11):247. TL. Pnom-Penh, Cambodia. (MNHN)
Papilio helenus aspadantus Fruhstorfer, [1916] : Arch. Naturgesch. (A)81(11):75. TL. Malayische Halbinsel. (NHML)


Type Materials Information

aulus ; ♂, Type (red), #149316, Hainan, Whitehead. (NHML, examined)
aspadantus ; ♂, Type (red), #149317, Malaka, H.Fruhstorfer (NHML, examined)
aurea ; Syntype 1♂, Pnom-Penh, Cambodge. (MNHN.)


Records taken from Literature

[ THAILAND ]
Papilio helenus helenus ; Godfrey,1916 : 110.
Papilio helenus helenus ; Pendlebury,1923 : 23-24. (Nakhon Sri Thammarat: Khao Luang; Tai Sai River)
Papilio helenus helenus ; Godfrey,1927 : 120. (N)
Papilio helenus helenus ; Godfrey,1930 : 212. (Prae; Muak Lek; Chonburi: Sriracha; Chiang Rai; Nakhon Sri Thammarat)
Papilio helenus helenus ; Godfrey,1932a : 264. (P)
Papilio helenus helenus ; Pinratana,1977 : 36, figs.P25,♂,♂(Un). (all over)
Papilio helenus helenus ; Pinratana & Eliot,1992 : 22,42 pl.28, figs.25,♂,♀, pl.29, figs.25,♂(Un),♂(ab.Un). (Chiang Mai; Chaiyaphum; Chanthaburi; Chonburi; Kanchanaburi; Nakhon Nayok; Ratchaburi; Ranong)
Papilio helenus helenus ; Katsuyama et al.,2006 : 46, fig.27,♂. (Nan: Ban Nam Poon)
Papilio helenus helenus ; Ek-Amnuay,[2007] : 78, pl.10, figs.P26,♂,♀. (Kanchanaburi)
Papilio helenus helenus ; Les Day,2011- : Samui Butterflies, figs. (Koh Samui) (accessed: 2022.01.19)
Papilio helenus helenus ; Kimura et al.,2011 : 123, figs.♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok; Doi Chiang Dao; Doi Suthep; Doi Inthanon; Thung Saleng Luang; Nam Nao; Phu Khiewo; Khao Yai; Nakohn Nayork; Wan Nam Khiewo; Kao Soi Dao; Kratchin WF; Khao Khiewo; Doi Musso; Sai York Noi; Hua Hin; Muang Shone; Khlong Naka; Ka Chong; Than To WF.)
Papilio helenus helenus ; Ek-Amnuay,2012 : 78, pl.10, figs.P24,♂,♀. (Kanchanaburi)
Papilio helenus helenus ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Papilio helenus helenus ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Phetchabun; Saraburi) (accessed: 2022.01.19)
Papilio helenus helenus ; Küppers & Janikorn,2017 : 158-159, figs.Abb.Pa-082-085,194-205. (Phang Nga)
Papilio helenus helenus ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Papilio helenus helenus ; Motono & Negishi,1989 : 46 pl.5 figs.5,♂,6,♂(Un),7,♀,8,♀(Un). (everywhere)
Papilio helenus helenus ; Osada, Uémura & Uehara,1999 : 200, pl.11, figs.♂(dsf),♂(dsf,Un),♀(dsf). (Lak Sao; Nong Het)
Papilio helenus helenus ; Nimura & Wakahara,2005 : 38. (Thabok; Nakai)
Papilio helenus helenus ; Cotton & Racheli,[2007] : 349-350. (throughout Laos)
Papilio helenus helenus ; Onodera,2009b : 36-37. (Xieng Kouang: Nam Noen)
Papilio helenus helenus ; Nakamura & Wakahara,2012 : 41.

[ CAMBODIA ]
Papilio helenus ab.aurea ; Dubois & Vitalis,1914 : 149. (Cambodge)
Papilio helenus ab.aurea ; Dubois & Vitalis,1919 : 213. (Cambodge)
Papilio helenus ab.aurea ; Dubois & Vitalis,1921 : 12. (Cambodge)
Papilio helenus helenus ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)

[ VIETNAM ]
Papilio helenus ; Joannis,1901 : 333. (Tonkin: Cao Bang)
Papilio helenus ; Fruhstorfer,[1903] : 173. (Tonkin: Chiem-Hoa / Annam: Phuc-Son; Xom-Gom)
Papilio helenus helenus ; Jordan,1908-1909 : 54. (Tonkin)
Papilio helenus ; Lemee,1950 : 4. (Hagiang; Hanoi)
Papilio helenus helenus ; Metaye,1957 : 82 (N / C / S)
Papilio helenus daksha ; Metaye,1957 : 82 (N / C / S)
Papilio helenus helenus ; Ikeda et al.,1998 : 17, figs.3:15,♂,16,♂(Un),4:1,♀,9:6(♂genitalia). (Cuc Phuong)
Papilio helenus helenus ; Shinkai,1999 : 53. (N: Tam Dao / S: Bao Lac)
Papilio helenus helenus ; Luong et al.,2004 : 1, 23, figs.♂,♂(Un). (Cuc Phuong)
Papilio helenus helenus ; Miyazaki, Saito & Saito,2006 : 7, figs.Pa-13,♂,♀. (Lam Dong: Dambri; Gung Re; Madagui)
Papilio helenus ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Papilio helenus helenus ; Monastyrskii,2007 : 55-56, pl.13, figs.2a♂,2b♂(Un),2c♀. (throughout Vietnam)
Papilio helenus helenus ; Monastyrskii & Devyatkin,2015 : 12. (N / C / S)
Papilio helenus helenus ; Yokochi et al.,2015 : 39. (Khanh Hoa: Hon Ba)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 10♂ 2♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 4♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 4♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Uttaradit : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 2♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 3♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Trat : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Ranong : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phuket : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Si Thammarat : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Yala : 3♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Xaignabouli : 7♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 27♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 20♂ 6♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Champasak : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 9♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is very common in montane and secondary forests at low to high elevations (alt.30m - 1600m). Males form flight paths along streams or forest edges, and frequently come to the banks of streams and puddles to suck water.


Literature

Linnaeus, C.,1758 : Systema naturae per regna tria naturae,secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis.
Holmiae, Laurentil Salvii. (Edition 10). 1(Animalia):[iv]+824pp.

Rothschild, L.W.,1895 : A revision of the Papilios of the Eastern Hemisphere, exclusive of Africa.
Novit. zool. 2(3):167-463, pl.4.

Fruhstorfer, H.,1908 : Neue ostasiatische Rhopaloceren.
Ent. Wochenbl. 25:38.

Boullet, E. & Le Cerf, F.,1912 : Descriptions sommarires de formes nouvelles de Papilionidae (Lep).
de la collection du Museum de Paris (2e note).
Bull. Soc. ent. Fr. 1912(11):246-247.

Fruhstorfer, H.,[1916] : Rhopaloceren aus Hollandisch-Neu-Guinea.
Arch. Naturgesch. (A)81(11):61-78.


Papilio helenus helenus ♂ Papilio helenus helenus ♀ with ♂
♂, Chiang Mai, Thailand. ♀ with ♂, Yala, Thailand.
Papilio helenus helenus ♂ Papilio helenus helenus ♂
2♂, Lampang, Thailand. ♂, Chiang Mai, Thailand.

1996.10.02 - 2022.03.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.