♂, Chiang Mai, Thailand.		♀, Phrae, Thailand.
♂, Phrae, Thailand.		♀, Chanthaburi, Thailand.
♂, Chiang Mai, Thailand.		♀, Nan, Thailand.

ON : Itanus phemius Doubleday, [1848] OD : Gen. diurn. Lep. (2): pl.41, fig.4,♂. TL : "Silhet" [southern Khasi Hills, India ; north of Sylhet, Bangladesh]. (NHML) Distribution : Nepal, Sikkim, Assam to Myanmar, S.E.Tibet, Yunnan, Thailand, Laos, Cambodia, Vietnam, Guangxi, Hainan, Guangdong. ● ssp.phemius ● ssp.ipona

Adolias sancara Moore, [1858]; In: Horsfield & Moore, Cat. lep. Ins. East-India Co. 1: 195, a♂[recte ♀], b♀; TL: 'N.India; Darjeeling [Darjeeling, W.Bengal, India]'. (NHML)
Euthalia phemius seitzi Fruhstorfer, 1913; In: Seitz, Gross-Schmetterl. Erde 9: 675; TL: Hongkong. (NHML)

phemius : Syntype ♂, Silhet. (NHML) ; Yago et al.,2012 : 306, figs.1A♂,1B♂(Un).
sancara : Syntypes "♂" [recte ♀], Rh.10126, N.India (NHML), ♀, Rh10127, Darjeeling (NHML) ; Yago et al.,2012 : 322, figs.21[20]1A♀,21[20]B♀(Un),22[21]A♀,22[21]B♀(Un).
zeitzi : Syntypes ♂, Tai Po Kau, Hong Kong. (MNHN), ♀, Hong Kong. (MNHN) ; Yago et al.,2012 : 306, figs.23[22]A♂,23[22]B♂(Un),24[23]A♀,24[23]B♀(Un),

[ THAILAND ]
Euthalia phemius phemius: Godfrey,1916 : 131. (Chonburi: Sriracha; Prae: Me Song forest)
Euthalia phemius phemius: Godfrey,1930 : 293. (Prae: Me Song forest; Chonburi: Sriracha)
Euthalia phemius phemius: Pinratana,1979 : 76, figs.N131,♂,♀,♀(Un). (Chiang Mai; Chanthaburi; Nakhon Ratchasima)
Euthalia phemius phemius: Pinratana & Eliot,1996 : 78, pl.53, figs.143,♂,♀. (Chiang Mai; Chanthaburi; Nakhon Ratchasima)
Euthalia phemius phemius: Ek-Amnuay,[2007] : 442, pl.195, figs.N148,♂,♂(Un),♀. (Chiang Mai)
Euthalia phemius phemius: Ek-Amnuay,2012 : 452, pl.199, figs.N314,♂,♂(Un),♀. (Chiang Mai)
Euthalia phemius phemius: Sophonviwatkul et al.,2015– : Butterflies of Thailand, figs. (Phetchabun; Sa Kaeo) (accessed: 2024.05.13)
Euthalia phemius phemius: Kimura et al.,2016 : 107, figs.♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Pha Hom Pok; Maethro; Bang Salok; Wan Nam Khiewo; Kao Soi Dao; Kratching WF.) (partim)

[ LAOS ]
Euthalia phemius phemius: Fruhstorfer,1913 : 674. (Lao States)
Euthalia phemius f. phemius: Dubois & Vitalis,1924 : 43. (Giranville)
Euthalia phemius phemius: Motono & Negishi,1989 : 60, Not fig. (Vang Vieng)
Euthalia phemius phemius: Osada et al.,1999 : 207, pl.64, figs.♂,♂,♀. (Nhahin; Phong Saly)
Euthalia phemius phemius: Nakamura & Wakahara,2012 : 46.
Euthalia phemius phemius: Onodera,2022 : 46, pl.133, figs.♂,♂(Un). (Oudomxai)

[ CAMBODIA ]
Euthalia phemius phemius: Kosterin,2019 : 97, fig.48,♀. (Kep)
Euthalia phemius phemius: Chartier & Kosterin,2022 : 118. (Kep; Koh Kong; Mondulkiri)

[ VIETNAM ]
Euthalia phemius phemius: Fruhstorfer,1913 : 674. (Tonkin)
Euthalia phemius f. phemius: Dubois & Vitalis,1924 : 43. (Tonkin: Vietri)
Euthalia phemius: Lemée,1950 : 15. (Hagiang)
Euthalia phemius phemius: Metaye,1957 : 97. (N)
Euthalia phemius phemius: Ikeda et al.,2000 : 36, figs.7:1,♀,2,♀(Un). (Cuc Phuong)
Euthalia phemius: Luong et al.,2004 : 7, 68, figs.♀,♀(Un). (Cuc Phuong)
Euthalia phemius: Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong; A Luoi; Bach Ma)
Euthalia phemius phemius: Miyazaki et al.,2006b : 32, figs.N-91,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Dong Nai: Thac Mai)
Euthalia phemius phemius: Monastyrskii & Devyatkin,2015 : 40. (N; C; S)
Euthalia phemius phemius: Monastyrskii,2019 : 4(2):120–121, pl.65, figs.1a♂,1b♀. (N: Ba Be; Tam Dao; Cuc Phuong; Pu Luong; C: Dakrong; Nam Dong; A Luoi; Bach Ma; Ba Na; Ba To; Phu Yen; Ninh Thauan; S: Nam Cat Tien; La Nga; Nghia Trung)
Euthalia phemius phemius: Saito et al.,2021 : 464. (Phu Quoc)

[ THAILAND ]
Chiang Mai : 15♂ 5♀ (incl. photo).

Jan

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Phrae : 1♂ 1♀.

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Nan : 8♂ 8♀.

Jan

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Loei : 1♂ (photo).

Jan

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Sakon Nakhon : 1♀ (photo by A.Giudici).

Jan

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Tak : 1♂ 1♀.

Jan

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Chaiyaphum : 1♂ (photo by A.Giudici).

Jan

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Sa Kaeo : 2♂ 1♀.

Jan

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Nakhon Nayok : 1♂ 1♀ (incl. photo).

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Chanthaburi : 5♂ 3♀.

Jan

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Trat : 1♂ 1♀.

Jan

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[ VIETNAM ]
Lam Dong : 3♂ 2♀.

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The butterfly is locally common in forests at low to high elevations (10–1,800 m a.s.l.).
Yago et al. (2012) identified ipona as a form of the nominotypical subspecies based on both morphology and molecular analysis. However, I treat ipona as a distinct subspecies, mainly on the basis of morphology.

Doubleday, E, [1848]. The genera of diurnal Lepidoptera: comprising their generic characters, a notice of their habits and transformations, and a catalogue of the species of each genus.
Longman, Brown, Green and Longman's, Lodon, 2: pls. 31–42. ([3.Nov.1848])

Moore, F, [1858]. In: Horsfield, T. & F. Moore, A catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company. Vol.1.
W.H. Allen and Co., London, v + 278 pp., 12 pls.

Fruhstorfer, H, 1913. 6. Familie: Nymphalidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde.
Alfred Kernen, Stuttgart, 9: 665–680. (13.Jan.1913)

Lang, S-Y, 2012. The Nymphalidae of China (Lepidoptera, Rhopalocera) Part 1.
Libytheinae, Danainae, Calinaginae, Morphinae, Heliconiinae, Nymphalinae, Charaxinae, Apaturinae, Cyrestinae, Biblidinae, Limenitinae.
Tshikolovets Publications, Pardubice, 456 pp., 32 col. + 41 b&w pls.

Yago, M, Yokochi, T, Kondo, M, Braby, M.F, Yahya, B, Peggie, D, Wang, M, Williams, M, Morita, S. & Ueshima, R, 2012.
Revision of the Euthalia phemius complex (Lepidoptera: Nymphalidae) based on morphology and molecular analyses.
Zoological Journal of the Linnean Society, 164: 204–324.

♂, Chiang Mai, Thailand.	♀, Chiang Mai, Thailand.
♂, Loei, Thailand.	♀, Nakhon Nayok, Thailand.
♂, Chaiyaphum, Thailand. (photo by A.Giudici)	♀, Sakon Nakhon, Thailand. (photo by A.Giudici)

2007.02.10 - 2026.03.31.