Cynitia cocytus ambrysus (Fruhstorfer, 1913)

Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un. Cynitia cocytus ambrysus ♀ Up. Cynitia cocytus ambrysus ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un. Cynitia cocytus ambrysus ♀ Up. Cynitia cocytus ambrysus ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un. Cynitia cocytus ambrysus ♀ Up. Cynitia cocytus ambrysus ♀ Un.
♂, Chanthaburi, Thailand. ♀, Chanthaburi, Thailand.
Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un. Cynitia cocytus ambrysus ♀ Up. Cynitia cocytus ambrysus ♀ Un.
♂, Chaiyaphum, Thailand. ♀, Chaiyaphum, Thailand.
Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un. Cynitia cocytus ambrysus ♂ Up. Cynitia cocytus ambrysus ♂ Un.
♂, Ninh Binh, Vietnam. ♂, Chanthaburi, Thailand.

ON : Euthalia cocytus ambrysus Fruhstorfer, 1913
OD : in Seitz, Grossschmett Erde 9:658, pl.132d. (as cocytus)
TL : Than Moi, "Nord-Tonkin" [N.Vietnam].
Distribution : Thailand, Laos, Cambodia, Vietnam, Yunnan, Guangxi.
ssp.cocytus
● ssp.ambrysus
ssp.aoyamai
Cynitia cocytus ambrysus map

Records taken from Literature

[ THAILAND ]
Euthalia cocytus cocytus ; Fruhstorfer,1913 : 658. (Siam)
Euthalia cocytus cocytus ; Godfrey,1916 : 130.
Euthalia cocytus cocytus ; Godfrey,1930 : 289-290. (Prae: Me Song forest; Muak Lek; Chonburi: Sriracha)
Euthalia cocytus cocytus ; Pinratana,1979 : 70, figs.N120,♂,♀,♀(Un). (Chaiyaphum; Chanthauburi; Kao Yai; Nakhon Ratchasima)
Tanaecia cocytus cocytus ; Pinratana & Eliot,1996 : 72, pl.47, figs.131,♂,♂(Un),♀. (Chiang Mai; Chaiyaphum; Nakhon Ratchasima; Nakhon Nayok; Chanthauburi) (in part)
Cynitia cocytus ambrysus ; Yamaguchi & Aoki,2002 : 29, fig.78,♂. (Ubon Ratchathani: Phu Chong Na Yoi; Kaeng E-Keow; Phu Hin Dang; Mukdahan: Ban Na Lak)
Cynitia cocytus ambrysus ; Yamaguchi, Aoki & Akiyama,2005 : 51. (Ubon Ratchathani: Nam Tok Soi Sawan; Phu Chong Na Yoi)
Tanaecia cocytus ambrysus ; Ek-Amnuay,[2007] : 426, pl.187, figs.N136,♂,♂(Un),♀,♀(Un). (Trat)
Cynitia cocytus cocytus ; Ek-Amnuay,2012 : 436, pl.191, figs.N302a♂,♀(Wiang Pa Pao),♀(Un,Wiang Pa Pao). (Chiang Mai) (in part)
Cynitia cocytus ambrysus ; Ek-Amnuay,2012 : 436, pl.191, figs.N302b♂,♀,♀(Un). (Chanthaburi; Trat)
Cynitia cocytus cocytus ; Kimura et al.,2016 : 101, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Chiang Dao; Doi Phui; Doi Suthep; Charn Phui; Phu Khiewo; Kao Yai; Sarika; Wan Nam Khiewo)
Cynitia cocytus ambrysus ; Kimura et al.,2016 : 101, figs.♂,♂(Un). (Kao Soi Dao; Kao Khiewo)
Cynitia cocytus ambrysus ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2024- : Butterflies of Thailand, figs. (Chiang Mai; Phetchabun; Khon Kaen; Sa Kaeo) (accessed: 2024.05.13)

[ LAOS ]
Euthalia cocytus f. satropaces ; Dubois & Vitalis,1924 : 42. (Laos)
Euthalia cocytus f. cocytus ; Dubois & Vitalis,1924 : 42. (Laos)
Tanaecia cocytus cocytus ; Motono & Negishi,1989 : 59, pl.66, figs.3,♂,4,♂(Un),5,♀,6,♀(Un). (Vang Vieng; Vientiane)
Cynitia cocytus ambrysus ; Osada, Uémura & Uehara,1999 : 207, pl.62, figs.♂,♀. (Lak Sao; Thakhek)
Cynitia cocytus ambrysus ; Nimura & Wakahara,2005 : 43. (Namdhua; Thabeng; Thakhek)
Cynitia cocytus ambrysus ; Nakamura & Wakahara,2012 : 46.
Cynitia cocytus ambrysus ; Suwa et al.,2013 : 18. (Petka; Nahin; Xebangfay; Thakhek; Thabok; Vang Vieng)
Cynitia cocytus ; Suwa et al.,2019 : 18. (Savannakhet; Dong Hua Sao; Thabok)
Cynitia cocytus ambrysus ; Onodera,2022 : 42, pl.122, figs.♂,♂(Un),♀,♀(Un),♀,♀(Un). (Savannakhet; Bolikhamxai: Phon Si)

[ CAMBODIA ]
Euthalia cocytus ; Vitalis,1919 : 231. (Cambodge)
Tanaecia cocytus cocytus ; Woodfield & Murton,2006 : fig,♀. (Botum Sakor NP.)
Cynitia cocytus ; Onodera,2007 : 28, fig.66,♂. (Siem Reap: Phnum Kuhlen)
Cynitia cocytus ; Onodera,2009 : 40. (Siem Reap: Phnum Kulen)
Cynitia cocytus ambrysus ; Chartier & Kosterin,2022 : 118. (Koh Kong; Mondulkiri; Siem Reap)

[ VIETNAM ]
Euthalia ambrysus ; Vitalis,1919 : 231. (Tonkin)
Euthalia cocytus f. ambrysus ; Dubois & Vitalis,1924 : 43. (Cochinchine: Gyaray)
Euthalia cocytus cocytus ; Metaye,1957 : 97. (N)
Euthalia cocytus ambrysus ; Metaye,1957 : 97. (S)
Tanaecia cocytus ; Luong et al.,2004 : 7, 66, figs.♀,♀(Un). (Cuc Phuong)
Cynitia cocytus ambrysus ; Miyazaki, Saito & Saito,2006b : 26, figs.N-83,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Cynitia cocytus ambrysus ; Monastyrskii & Devyatkin,2015 : 39. (N / C / S)
Cynitia cocytus ambrysus ; Monastyrskii,2019 : 4(2):107-108, pl.60, figs.1a♂,1b♀. (N: Ba Be; Huu Lien; Cuc Phuong / C: Ben En; Pu Mat; Vu Quang; Phong Nha/Ke Bang; Da Teh; Dong Xuan ; Bai Ai; Bao Lam; Cat Loc / S: Cat Tien; Lo Go Sa Mat; Tan Phu; La Nga; Nghia Trung; Binh Chau)


Specimens Examined

[ THAILAND ]
Chiang Mai : 8♂ 7♀ (incl. photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 1♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Sa Kaeo : 2♂ (photo by M.Hayashi).
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Nayok : 1♂ 1♀ (photo & photo by M.Hayashi).
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 5♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Ninh Binh : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Lam Dong : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is common in montane forest at low to moderate elevations (Alt. 150 - 1200m) and males are territorial on shrubs or undergrowth.
The population in Chiang Mai occasionally includes individuals somewhat similar to ssp.cocytus. It seems a transition zone of both subspecies.


Literature

Fruhstorfer, H.,1913 : 6. Familie: Nymphalidae in Seitz, Gross-Schmett. Erde 9
657-664. (12.Jan.1913)


Cynitia cocytus ambrysus ♂ Cynitia cocytus ambrysus ♀
♂, Nakhon Nayok, Thailand. (photo by M.Hayashi) ♀, Nakhon Nayok, Thailand.
Cynitia cocytus ambrysus ♂ Cynitia cocytus ambrysus ♀
♂, Sa Kaeo, Thailand. (photo by M.Hayashi) ♀, Chiang Mai, Thailand.
Cynitia cocytus ambrysus ♂ Cynitia cocytus ambrysus ♀
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Cynitia cocytus ambrysus ♂ Cynitia cocytus ambrysus ♀
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.

2007.01.31 - 2024.05.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.