♂, Chanthaburi, Thailand.		♀, Chanthaburi, Thailand.
♂, Chiang Mai, Thailand.		♀, Chaiyaphum, Thailand.
♂, Ranong, Thailand.		♀, Ranong, Thailand.
♂, Tak, Thailand.		♀, Satun, Thailand.
♂, Bolikhamxai, Laos.		♂, Phong Saly, Laos.
♂, Nan (Pua dist.), Thailand.		♂ genitalia, Chiang Mai, Thailand.

ON : Neptis nolana H. Druce, 1874 OD : Proc. zool. Soc. Lond. 1874(1):105. TL : "Chentaboon, Siam" [Chanthaburi, Thailand]. (NHML) Distribution : Myanmar, Thailand, Laos, Cambodia, Vietnam, S.Yunnan, Fujian, Hainan. ● ssp.nolana ● ssp.thaithummakulorum ● ssp.disopa ● ssp.batara

Type ♀, "Chentaboon, Siam" [Chanthaburi, Thailand]. (NHML)

[ THAILAND ]
Neptis miah nolana ; Fruhstorfer,1912 : 621. (Siam)
Neptis miah nolana ; Godfrey,1916 : 129. (Pak Jong)
Neptis miah nolana ; Godfrey,1930 : 283. (Pak Jong; Muak Lek)
Neptis miah nolana ; Eliot,1969 : 89. (Siam)
Neptis miah nolana ; Pinratana,1979 : 48, figs.N81a♂,♂(Un). (Chiang Mai; Chaiyaphum; Chanthaburi; Nakhon Nayok; Nakhon Ratchasima; Prachinburi)
Neptis miah nolana ; Pinratana & Eliot,1996 : 50, pl.30, figs.74a♂,♂(Un). (Chiang Mai; Chaiyaphum; Nakhon Ratchasima; Nakhon Nayok; Prachinburi; Chanthaburi)
Neptis miah nolana ; Yamaguchi & Aoki,2002 : 27, fig.74,♀. (Mukdahan: Ban Na Lak)
Neptis miah nolana ; Yamaguchi, Aoki & Akiyama,2005 : 50. (Ubon Ratchathani: Phu Chong Na Yoi)
Neptis miah nolana ; Ek-Amnuay,[2007] : 390, pl.169, figs.N77a♂,♂(Un),♀[♂],♀(Un)[♂](Un). (Phu Khieo)
Neptis miah nolana ; Les Day,2011-2022 : Samui Butterflies, figs.♂,♂(Un). (Koh Samui) (accessed: 2022.06.10)
Neptis miah nolana ; Ek-Amnuay,2012 : 402,404, pl.174, figsN253a,♂,♂(Un),♀[♂],♀(Un)[♂](Un). (Phu Khieo)
Neptis miah nolana ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Nakhon Ratchasima; Phetchaburi) (accessed: 2023.06.13)
Neptis miah nolana ; Kimura et al.,2016 : 79, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Suthep; Doi Inthanon; Thung Saleng Luang; Phu Khiewo; Nakohn Nayok; Wan Nam Khiewo; Kao Soi Dao; Kratching WF; Kao Kiewo)
Neptis miah batara ; Küppers & Janikorn,2020 : 53-54, figs.Abb.N-297-299. (Phang Nga)
Neptis ananta learmondi ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2024- : Butterflies of Thailand, fig. (19/02/2015 Chiang Mai) (in part) (accessed: 2024.05.12)

[ LAOS ]
Neptis miah f. nolana ; Dubois & Vitalis,1924 : 40. (Ban-Sen-Savouane; Pou-So-Ko; Son-Hat)
Neptis miah nolana ; Motono & Negishi,1989 : 57, pl.57, figs.7,♂,8,♂(Un). (Vang Vieng; Tha Ngone; Luang Prabang)
Neptis miah nolana ; Osada, Uémura & Uehara,1999 : 206, pl.55, figs.♂,♂(Un),♀. (Nong Het; Vang Vien; Oudomxay)
Neptis miah nolana ; Nimura & Wakahara,2005 : 42. (Nakai)
Neptis miah nolana ; Nakamura & Wakahara,2012 : 45.
Neptis miah nolana ; Onodera,2022 : 35, pl.112, figs.♂,♂(Un),♀,♀(Un). (Vientiane: Vang Vieng; Xiengkhouang: Tha)

[ CAMBODIA ]
Neptis miah nolana ; Chartier & Kosterin,2022 : 117, fig.2H. (Siem Reap)

[ VIETNAM ]
Neptis miah nolana ; Fruhstorfer,1912 : 621. (Annam) (in part)
Neptis miah nolana ; Godfrey,1919 : 468. (S: Daban)
Neptis nolana ; Vitalis,1919 : 230. (Annam) (in part)
Neptis miah f. nolana ; Dubois & Vitalis,1924 : 40. (Annam) (in part)
Neptis miah nolana ; Metaye,1957 : 96. (C) (in part)
Neptis miah ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien)
Neptis miah nolana ; Miyazaki, Saito & Saito,2006b : 22, figs.N-55,♂,♀. (Lam Dong: Dambri; Dong Nai: Thac Mai)
Neptis miah nolana ; Yokochi et al.,2015 : 40. (Khanh Hoa: Hon Ba)
Neptis miah ; Monastyrskii & Devyatkin,2015 : 37. (C / S) (in part)
Neptis miah nolana ; Monastyrskii,2019 : 4(2):70-72, pl.46, figs.1d♂,1e♀,1f♀(Un). (C: Pu Hoat; Pu Mat; Huong Son; Vu Quang; Phong Dien; Nam Dong; A Luoi; Huong Thuy; Ba Na; Kon Ka Kinh; Ba To; Chu Yang Sin; Hon Ba; Da Teh; Bao Lam; Son Thanh / S: Cat Tien; La Nga)

[ THAILAND ]
Mae Hong Son : 3♂.

Jan

Feb

Mar

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May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Rai : 1♂ (photo).

Jan

Feb

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Chiang Mai : 6♂ 1♀.

Jan

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Nan (Chiang Klang dist ; Pua dist.) : 2♂ 1♀ (incl photo by A.Giudici).
Pua dist., 2020.03.23. 1♂ A.Giudici (YI).

Jan

Feb

Mar

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May

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Oct

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Dec

Tak : 3♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

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Dec

Chaiyaphum : 2♂ 3♀ (incl. photo).

Jan

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May

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Jul

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Sep

Oct

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Chanthaburi : 6♂ 5♀.

Jan

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Oct

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Ranong : 3♀.

Jan

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Mar

Apr

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Oct

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Surat Thani : 1♀.

Jan

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May

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Satun : 1♀.

Jan

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Dec

[ LAOS ]
Bolikhamxai : 3♂ (incl. photo by H.Onodera).

Jan

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Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phong Saly : 1♂.

Jan

Feb

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Vientiane : 1♂ (photo by H.Onodera).

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The butterfly is common in montane forests at low to moderate elevations (Alt. 50 - 1300m) and is frequently found on forest paths or forest edges.
In Pua dist., Nan, hitherto only ssp.thaithummakulorum had been known for over 30 years, and ssp.nolana is found only in Chiang Klang dist., the opposite mountain range to Pua dist.
However, recently in 2020 - 2021, We found ssp.thaithummakulorum and ssp.nolana occurring together in Pua dist. Possibly, ssp.nolana might be expanding its distribution due to environmental changes, such as desiccation. Currently, both subspecies are distinct and there are no intermediate individuals. It will be very interesting to see how the relationship between these two subspecies will progress.

Druce, H.,1874 : A List of the Lepidopterous Insects collected by Mr. L. Layard at Chentaboon and Nahconchaisee, Siam,
with Descriptions of new Species.
Proc. zool. Soc. Lond. 1874(1):102-109, pl.16, 6 figs.

Eliot, J.N.,1969 : An Analysis of the Eurasian and Australian Neptini (Lepidoptera: Nymphalidae).
Bull. Br. Mus. nat. Hist. (Ent.) Supple.15. London.

♂, Chiang Rai, Thailand.	♂, Chiang Mai, Thailand.
♂, Chaiyaphum, Thailand.	♀, Nan (Chiang Klang dist.), Thailand. (photo by A.Giudici)
♂, Nan (Pua dist.), Thailand.	♂, Nan (Pua dist.), Thailand. (photo by A.Giudici)

2006.02.20 - 2024.05.31.