♂, Surat Thani, Thailand.		♀, 2010.04.06. Surat Thani, Thailand. (photo by L.Day)
♂, Kien Giang, Vietnam. (photo by E.Ikari)		♀, Kien Giang, Vietnam. (photo by E.Ikari)

ON : Ismene leucospila Mabille, 1891 OD : C. R. Soc. ent. Belg. 35:lxxix. TL : Minah. [Minahassa, N.Sulawesi]. (♂) Distribution : S.Myanmar, P.Thailand, Cambodia, S.Vietnam, W.Malaysia, Sumatra, Java, Borneo, Philippines, Sulawesi.

Hasora palinda Swinhoe, 1905 : Ann. Mag. nat. Hist. (7)16:618. TL. Java. (♂, NHML)
Hasora leucospila matisca Fruhstorfer, 1911 : Deut. ent. Zeit. [Iris] 25:76. TL. Bazilan. (♂, NHML)
Hasora leucospila parnia Fruhstorfer, 1911 : Deut. ent. Zeit. [Iris] 25:76. TL. Kina Balu, N.Borneo. (♂, NHML)
Hasora borneensis mavis Evans, 1934 : Entomologist 67:34. TL. Perak, Malaysia. (♀, NHML)

[ THAILAND ]
Hasora leucospila leucospila ; Pinratana,1985 : 21, pl.7, fig.21,♂(Un). (Ranong; Surat Thani)
Hasora mavis ; Maruyama,1991 : 7. (Ko Samui)
Hasora leucospila leucospila ; Kimura,1996 : 21, pl.3, figs.41,♂,42,♂(Un). (Surat Thani: Kao Poh Ta)
Hasora leocospila leucospila ; Ek-Amnuay,[2007] : 736, pl.337, figs.H25,♂,♂(Un). (Ranong)
'Hasora mavis ; Ek-Amnuay,[2007] : 738, pl.338, fig.H27,♀(HT, Malaysia).'
Hasora leucospila leucospila ; Les Day,2011-2022 : Samui Butterflies, fig.♂(Un). (Koh Samui) (accessed: 2023.03.08)
Hasora mavis ; Les Day,2011-2022 : Samui Butterflies, figs.♂[♀](Un),♀(Un). (Koh Samui) (accessed: 2023.03.08)
Hasora mavis ; Kimura et al.,2011 : 27. (by literature)
Hasora leucospila leucospila ; Kimura et al.,2011 : 28, figs.♂,♂(Un). (Muang Shone; Khlong Naka)
Hasora leucospila leucospila ; Ek-Amnuay,2012 : 784, pl.361, figs.H24,♂,♂(Un). (Ranong)
Hasora mavis ; Ek-Amnuay,2012 : 786, pl.362, figs.H26,♀,♀(Un). (Koh Samui)

[ CAMBODIA ]
Hasora leucospila leucospila ; Chartier & Kosterin,2022 : 124, fig.6I♂. (Koh Kong)

[ VIETNAM ]
Hasora mavis ; Sugimoto,2019 : 52, figs.1,♀,♀(Un). (Kien Giang: Phu Quoc)
Hasora leucospila ; Sugimoto,2019 : 52, figs.2,♂,♂(Un). (Kien Giang: Phu Quoc)
Hasora leucospila leucospila ; Xue et al.,2019 : 391, figs.1a♀,1b♀(Un),2a♂,2b♂(Un). (Kien Giang: Phu Quoc)

[ THAILAND ]
Surat Thani (Koh Samui dist ; Koh Phangan dist.) : 3♂ 1♀ (incl. photo by A.Giudici and K.Yano).

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[ VIETNAM ]
Kien Giang (Phu Quoc) : 8♂ 2♀.

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The butterfly is locally not rare in lowland forests (Alt. 10 - 200m) and is crepuscular. Males frequently come to the banks of streams and puddles to suck water.
Hasora mavis was known as a distinct species for a long time. However, it was confirmed as a female of Hasora leucospila by Xue et al.,2019.

Mabille, P.,1891 : Description d'Hesperides nouvelles. Premitere Partie, Deuxieme Partie, Troisieme Partie.
C. R. Soc. ent. Belg. 35:lix-lxxxviii, cvi-cxxi, clxviii-clxxxvii.

Swinhoe, C.,1905 : On new and little-known Species of Eastern and Australian Lepidoptera.
Ann. Mag. nat. Hist. (7)16:612-629.

Fruhstorfer, H.,1911 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 25(2-6):9-79, pl.1.

Evans, W.H.,1934 : Indo-Australian Hesperiidae: Description of new Genera, Species and Subspecies.
Entomologist 67(2):33-36 (part.1), (3):61-65 (part.2), (7):141-151 (part.3), (8):181-184 (part.4), (9): 206-209 (part.5), (10): 231-234 (part.6).

Maruyama, K.,1991 : Butterflies of Borneo. Vol.2, No.2. Hesperiidae.
Tobishima Corporation.

Kitamura, M.,2002 : Butterflies from the southwest side slope of Mt. Banahaw, Mid-south Luzon, Philippines (7) Hesperiidae. Part II, Satyridae.
Butterflies 34:43-57.

Sugimoto, S.,2019 : Notes on some Hesperid butterflies of Southeast Asia
Butterflies 80:52-54.

Xue, G., Ikari, E., Inayoshi, Y. & Li, M.,2019 :
Hasora mavis Evans,1934 syn.n. is confirmed to be the female of H.leucospila leucospila (Mabille,1891) (Hesperiidae, Coeliadinae) by DNA barcoding
Zootaxa 4695(4):391-393.

♂, Surat Thani, Thailand. (photo by A.Giudici)	Larva, Surat Thani, Thailand. (photo by K.Yano)

1999.07.31 - 2023.03.31.