♂, Chanthaburi, Thailand.		♀, Chachoengsao, Thailand. (photo by THNHM)
♂, Chiang Mai, Thailand.		♂, Tak, Thailand.
♂, Tak, Thailand.		♂, Tak, Thailand.
♂, Chiang Mai, Thailand.		♂, Chiang Mai, Thailand.
♂, Lam Dong, Vietnam. (photo by K.Saito)		♀, Lam Dong, Vietnam. (photo by K.Saito)

ON : Ismene jaina margana Fruhstorfer, 1911 OD : Deut. ent. Zeit. [Iris] 25(5): 60. TL : "Hinlap, Siam" [Hin Lap, Saraburi, Thailand]. (NHML) Distribution : Thailand, Laos, Cambodia, Vietnam. ● ssp.jaina ● transition zone of jaina and margana ● ssp.margana ● transition zone of margana and velva ● ssp.velva

♂ Type (red), H2386. Jan. Hinlap, Siam. H.Fruhstorfer (NHML, examined)

[ THAILAND ]
Ismene jaina morgana [sic] ; Godfrey,1916 : 146. (Pak Jong)
Ismene jaina margana ; Seitz,1927 : 1054. (Siam)
Ismene jaina margana ; Godfrey,1930 : 355. (Pak Jong; Hinlap)
Bibasis jaina margana ; Evans,1949 : 47. (Siam)
Bibasis jaina margana ; Pinratana,1985 : 14, pl.1, figs.2b♂,♂(Un). (Chaiyaphum)
Bibasis jaina velva ; Pinratana,1985 : 14, pl.2, fig.2c♂. (Chaiyaphum; Ratchaburi)
Burara jaina jaina ; Ek-Amnuay,[2007] : 720, pl.329, figs.H2a♂,♂(Un). (Chiang Mai; Khao Soi Dao)
Burara jaina margana ; Ek-Amnuay,[2007] : 720, pl.329, figs.H2b♂,♂(Un). (Phu Khieo; Ratchaburi)
Burara jaina margana ; Chiba,2009 : 9, pl.2, fig.3,♂. (Chiang Mai; N.Soidao; Chantaburi)
Burara jaina margana ; Kimura et al.,2011 : 21, figs.♂,♂(Un). (Phu Khiewo; Wan Nam Khiewo)
Burara jaina jaina ; Ek-Amnuay,2012 : 768, pl.353, figs.H2a♂,♂(Un),♂,♂(Un). (Chiang Mai; Khao Soi Dao; Phu Khieo; Ratchaburi)
Burara jaina margana ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Phetchaburi; Sa Kaeo) (accessed: 2022.12.08)

[ LAOS ]
Burara jaina margana ; Masui & Uehara,1999 : 19,22, figs.17b♂,♂(Un). (Thakhek)
Burara jaina margana ; Osada, Uémura & Uehara,1999 : 221, pl.130, fig.♂. (Thakhek)
Burara jaina margana ; Chiba,2009 : 9, pl.2, fig.3,♂. (Thakhek)
Burara jaina margana ; Nakamura & Wakahara,2012 : 56.

[ CAMBODIA ]
Burara jaina margana ; Chartier & Kosterin,2022 : 124, fig.6A. (Koh Kong)

[ VIETNAM ]
Bibasis jaina margana ; Devyatkin & Monastyrskii,1999 : 154. (C: Bach Ma)
Bibasis jaina margana ; Devyatkin & Monastyrskii,2003 : 121. (S: Cat Tien)
Bibasis jaina ; Monastyrskii et al.,2006 : 44. (Thua Thien Hue: Nam Dong; A Luoi; Phong Dien; Bach Ma)
Burara jaina margana ; Miyazaki, Saito & Saito,2007b : 2, fig.H-1,♂. (Lam Dong: Dai Duc Me)
Burara jaina margana ; Miyazaki, Saito & Saito,2007c : 40, fig.H-1,♀. (Lam Dong: Dai Duc Me)
Burara jaina margana ; Chiba,2009 : 9, pl.2, fig.3,♂. (C / S)
Bibasis jaina margana ; Monastyrskii & Devyatkin,2015 : 70. (C / S)

[ THAILAND ]
Chiang Mai : 8♂.

Jan

Feb

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Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 8♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

Oct

Nov

Dec

Nakhon Nayok : 1♂ (photo by T.Aoyama).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

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Dec

Chachoengsao : 1♀ (photo by THNHM).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Sa Kaeo : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

Oct

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Dec

Chanthaburi : 5♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

Oct

Nov

Dec

[ VIETNAM ]
Lam Dong : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

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Dec

The butterfly is locally not rare in montane forests at low to moderate elevations (Alt. 150 - 600m) and is crepuscular. Males frequently come to the banks of streams and puddles to suck water.
The populations from Tak to Chiang Mai have quite variable and unstable male sex brand on the UpF. Some specimens from Chiang Mai have a small brand like ssp.jaina and some specimens from Tak are close to ssp.velva. It seems to be transitional with intermediate individuals between each subspecies.

Fruhstorfer, H.,1911 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 25(5): 57-72. (1.May.1911)

Evans, W.H.,1949 : A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History).
i-xix + 1-502, pls.1-35. London.

♂, Sa Kaeo, Thailand.

2011.12.31 - 2023.02.28.