♂ (dsf), Lam Dong, Vietnam. (photo by TME)		♀ (dsf), Chiang Mai, Thailand.
♂ (dsf), Chiang Mai, Thailand.		♀ (dsf), Mae Hong Son, Thailand.
♂, Chiang Mai, Thailand.		♂ ab., Chiang Mai, Thailand.

ON : Papilio lisandra Cramer, 1780 OD : Uitl. Kapellen 4(25):11, pl.293, figs.G[♀],H[♀,Un]. TL : S.China. Distribution : Thailand, Laos, Cambodia, Vietnam, S.China, Hainan.

Yphthima [sic] micrommatus Holland, 1887 : Trans. amer. ent. Soc. 11:116, pl.2, fig.3,♂,♂(Un). TL. Hainan.
Ypthima baldus scota Fruhstorefer, 1911 : in Seitz, Grossschmett. Erde 9:289. TL. Hong Kong (♀, NHML).

[ THAILAND ]
Ypthima lisandra bara ; Aoki & Uémura,1984 : 82-83. figs.9i-j. (Mae Hong Son: Mae Hong Son; Chiang Mai: Tha Tong; Mae Sa; Doi Suthep; Ban Mae Wa; Nam Tok Mae Klang; Kanchanaburi: Kanchanaburi; Sai Yok Noi; Loei: Phu Kadung; Nam Nao; Kalasin: Ban Lam Pao)
Ypthima lisandra bara ; Pinratana,1988 : 32, pl.33, figs.87,♂,♀(Un). (Chiang Mai; Chaiyaphum; Nakhon Ratchasima; Petchaboon)
Ypthima lisandra bara ; Yamaguchi & Aoki,2000 : 253-254, fig.78,♂(Un). (Mukdahan: Phu Pha Thaep; Ban Naudom; Phu Huay Sing; Ubon Ratchathani: Pha Taem)
Ypthima lisandra bara ; Yamaguchi & Aoki,2002 : 24. (Mukdahan: Ban Lao Kram; Nam Tok Kham Cha I; Ban Na Lak; Ubon Ratchathani: Ban Ba Hai; Ban Na Pho Krang)
Ypthima lisandra bara ; Yamaguchi, Aoki & Akiyama,2005 : 49. (Mukdahan: Ban Lao Kram; Phu Hin Khan; Ubon Ratchathani: Kaen Tana; Ban Ba Hai)
Ypthima lisandra bara ; Katsuyama et al.,2005 : 68, fig.127,♂. (Umphang)
Ypthima lisandra bara ; Katsuyama et al.,2006 : 49, fig.99,♂. (Nan: Ban Nam Poon)
Ypthima lisandra bara ; Ek-Amnuay,[2007] : 298, pl.121, fig.S95,♂(Un). (Chiang Mai)
Ypthima lisandra bara ; Ek-Amnuay,2012 : 298, pl.121, fig.N130,♂(Un). (Chiang Mai)
Ypthima lisandra bara ; Kimura et al.,2016 : 200, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Doi Suthep; Thoen; Wan Nam Khiewo; Chaiya Phoong; Ban Lao Kram; Bang Phra; Sai York Noi; Kaeng Chai)
Ypthima lisandra lisandra ; Uémura,2022 : 4-6. (Mae Hong Son; Tha Tong; Nam Tok Mae Sa; Doi Suthep; Ban Mae Wa; Nam Tok Mae Klang; Sai Yok Noi; Kanchanaburi; Phu Kadung; Nam Nao; Ban Lam Pao)
Ypthima lisandra lisandra ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2024- : Butterflies of Thailand, figs. (Phetchabun) (accessed: 2024.05.06)

[ LAOS ]
Ypthima lisandra f. lisandra ; Dubois & Vitalis,1924 : 27. (Ban-Pou)
Ypthima lisandra bara ; Osada, Uémura & Uehara,1999 : 211, pl.96, figs.♂,♂(Un). (Lak Sao; Xiang Khouang)
Ypthima lisandra bara ; Uémura,2005 : 2, fig.11 map. (Xaignabouri; Xiang Khouang; Lak Sao)
Ypthima lisandra bara ; Uémura,2011 : 4. (C: Nakhai to Lak Sao; Phin-Tat Hai)
Ypthima lisandra bara ; Nakamura & Wakahara,2012 : 49.
Ypthima lisandra lisandra ; Uémura,2022 : 4-6. (Xaignabouri: Mt Phaxang; Xiang Khouang; Lak Sao; Nakhai to Lak Sao; Phin-Tat Hai)
Ypthima lisandra lisandra ; Onodera,2022 : 62, pl.183, figs.♂(wsf),♂(wsf,Un),♀(dsf),♀(dsf,Un),♀(dsf),♀(dsf,Un). (Xiengkhouang: Plain of Jars)

[ CAMBODIA ]
Ypthima lisandra lisandra ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)
Ypthima lisandra lisandra ; Chartier & Kosterin,2022 : 116. (Pursat)

[ VIETNAM ]
Ypthima lisandra lisandra ; Fruhstorfer,1911 : 288. (Tonkin: Manson Mts.)
Ipthima [sic] lisandra ; Vitalis,1919 : 222. (Tonkin)
Ypthima lisandra f. lisandra ; Dubois & Vitalis,1924 : 27. (Tonkin)
Ypthima lisandra lisandra ; Metaye,1957 : 102. (N)
Ypthima lisandra lisandra ; Uémura & Monastyrskii,2004 : 28-29, figs.2x, 6.6,♂(wsf), 6.9,♂(dsf). (C: Play Ku; Nui Chua; Yok Don; Ban Don; Da Lat / S: Lo Go Sa Mat)
Ypthima lisandra lisandra ; Monastyrskii,2005 : 121-122, pl.32, figs.2a♂(dsf),2b♂(dsf,Un),2c♂(wsf),2d♂(wsf,Un). (N: 'Mau Son Mountains' / C: Play Ku; Nui Chua; Yok Don; Ban Don; Da Lat; Bao Lam / S: Lo Go Sa Mat)
Ypthima lisandra lisandra ; Miyazaki et al.,2010 : 24, fig.S-61,♀. (Lam Dong: Gung Re)
Ypthima lisandra lisandra ; Monastyrskii & Devyatkin,2015 : 26. (C / S)
Ypthima lisandra lisandra ; Uémura,2022 : 4-6. (C: Gia Lai: Play Ku; Ninh Thuan: Nui Chua; Dac Lac: Yok Don; Ban Don; Lam Dong: Da Lat; Bao Lam / S: Tay Ninh: Lo Go Sa Mat)

[ THAILAND ]
Mae Hong Son : 3♂ 2♀.

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Chiang Mai : 13♂ 8♀ (incl. photo).

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Loei : 2♂ 1♀.

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[ LAOS ]
Xieng Khouang : 1♂ 2♀ (photo by H.Onodera).

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[ VIETNAM ]
Lam Dong : 1♂ (photo by TME).

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The butterfly is locally common in semi-dried forests and grasslands at low to moderate elevations (Alt. 250 - 1100m).
Uémura,2022 treated the taxon bara as a subspecies of avantha. The populations from Thailand and Laos, previously thought to be lisandra bara, were then treated as the nominotypical subspecies.

Cramer, P.,1775-1780 :
De uitlandsche Kapellen voorkomende in de drie Waereld-Deelen Asia, Africa en America. Papillons exotiques des trois parties du monde l'Asie, l'Afrique et l'Amerique.
Amsteldam, S.J.Baalde; Barthelemy Wild and J. Van Schoonhoven & Comp.
1(1/7):i-xxx, 1-16, 1-132, pls.1-84 ([31 December]1775), (8):133-155, pls.85-96 (1776); 2(9/16):1-151, pls.97-192 (1777); 3(17/22):1-128, pls.193-264 (1779), (23/24):129-176, pls.265-288 (1780); 4(25/26):1-28, pls.289-304 (1780)

Holland, W.J.,1887 : Notes upon a small collection of Rhopalocera made by Rev. B. C. Henry in the Island of Hainan,
together with descriptions of some apparently new species.
Trans. amer. ent. Soc. 14:111-124, 2 pls.

Fruhstorfer, H.,1911 : 4. Familie: Satyridae in Seitz, Gross-Schmett. Erde 9
289-296. (15.Feb.1911)

Uémura, Y.,2022 : A revisional guide to the philomela-group of the genus Ypthima Hübner (Lepidoptera: Nymphalidae, Satyrinae) Part II
Bull. Toyosato Mus. Ent. (31):1-12.

♂ (dsf), Chiang Mai, Thailand.	♀, Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.	♀, Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.

1998.12.08 - 2024.05.31.