♂ (dsf), Mae Hong Son, Thailand.		♀ (dsf), Chiang Mai, Thailand.
♂, Nan, Thailand.		♀, Mae Hong Son, Thailand.
♂ (wsf), Mae Hong Son, Thailand.		♂, Ninh Binh, Vietnam.
♂, Yala, Thailand.		♂, Yala, Thailand.

ON : Appias dapha Moore, 1879 OD : Proc. zool. Soc. Lond. 1878(4): 838-839. TL : "Moulmein to Meetan" [Mawlamyine, Mon State to Kayin State, Myanmar]. (NHML) Distribution : Myanmar, C. to W.Yunnan, Thailand, Laos, Cambodia, Vietnam, Langkawi, W.Malaysia (Kedah). ● ssp.dapha ● ssp.nerissa

Cepora nerissa dapha wet-season form depuncta Talbot, 1939 : Fauna Brit. India (Butts) (ed.2) 1: 366. TL. "Bhamo district, Burma" [Bhamo Dist., Kachin State, Myanmar]. (NHML) (IFS)
Cepora nerissa yunnanensis Mell, 1943 : Zool. Stuttgart 36(100): 98. TL. "südlich Tali, Westyunnan" [Dali, W.Yunnan].

dapha ;
♂, Type (red), Moulmein to Meetan. (NHML, examined)
♀, Type (red), Moulmein to Meetan, Upper Tenasserim. (NHML, examined)
depuncta ;
♂, Type HT (red), 17.5.38. Bhamo Dist., E.Burma, 700ft., T.P.Dewar. (NHML, examined)
♀, Type AT (red), May-Aug.1928. East Bhamo Dist., Upper Burma. T.P.Dewar. (NHML, examined)

[ THAILAND ]
Huphina nerissa dapha ; Fruhstorfer,1910 : 141. (Siam)
Huphina nerissa dapha ; Godfrey,1916 : 114.
Huphina nerissa dapha ; Godfrey,1927 : 120. (N)
Huphina nerissa dapha ; Godfrey,1930 : 224. (all over)
Huphina nerissa dapha ; Godfrey,1931 : 192. (Ko Tao)
Huphina nerissa dapha ; Godfrey,1932a : 265. (P)
Cepora nerissa dapha ; Pinratana,1983 : 14, pl.12, figs.22,♀,♂(Un). (Chiang Mai; Chaiyaphum; Chanthaburi; Kanchanaburi; Loei; Nakhon Ratchasima; Phetchabun; Tak; Saraburi; Uthai Thani; Nonthaburi; Chonburi; Phetchaburi; Prachuap Khiri Khan; Trang)
Cepora nerissa dapha ; Yamaguchi, Aoki & Akiyama,2005 : 46, 58, fig.11,♂. (Mukdahan: Phu Hin Khan)
Cepora nerissa dapha ; Ek-Amnuay,[2007] : 144, pl.44, figs.Pi24,♂,♂(Un),♂(dsf),♂(dsf,Un),♀,♀(Un),♂(dsf),♂(dsf,Un). (Ban Hong; Wang Chin; Mae Hong Son)
Cepora nerissa daphne [sic] ; Les Day,2011-2022 : Samui Butterflies, figs.[♀],♂[♀](Un). (Koh Samui) (accessed: 2022.08.03)
Cepora nerissa dapha ; Kimura et al.,2011 : 162, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok; Chiang Dao; Thung Saleng Luang; Phu Kradung; Phu Khiewo; Sarika; Wan Nam Khiewo; Bang Phra; Kao Soi Dao; Sai York Noi; Kanchanaburi; Kaeng Krachan; Than To WF.)
Cepora nerissa dapha ; Ek-Amnuay,2012 : 144, pl.44, figs.Pi24,♂,♂(Un),♂(dsf),♂(dsf,Un),♀,♀(Un),♂(dsf),♂(dsf,Un). (Ban Hong; Wang Chin; Mae Hong Son)
Cepora nerissa dapha ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Cepora nerissa dapha ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Sa Kaeo; Bangkok; Saraburi; Phetchaburi) (accessed: 2024.04.18)
Cepora nerissa dapha ; Küppers & Janikorn,2017 : 192-193, figs.Abb.Pi-039-040. (Phang Nga)
Cepora nerissa dapha ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Huphina nerissa f. evagete ; Dubois & Vitalis,1921 : 19. (Laos)
Huphina nerissa f. zeuxippe ; Dubois & Vitalis,1921 : 19. (Laos)
Cepora nerissa dapha ; Motono & Negishi,1989 : 49, pl.18, figs.7,♂,8,♂(Un) ; pl.19, figs.1,♀(dsf),2,♀(dsf,Un),3,♀(wsf),4,♀(wsf,Un). (Pakse; Tha Ngone; Vientiane; Nam Moh; Luang Prabang; Don Dok; Tha Dua)
Cepora nerissa dapha ; Osada, Uémura & Uehara,1999 : 202, pl.33, figs.♂(dsf),♂(dsf,Un),♀(dsf),♂,♂(Un),♀. (Xam Neua; Vientiane; Nong Het; Luang Phrabang)
Cepora nerissa dapha ; Onodera,2009b : 34-35. (Vientiane: Namsong Riverside)
Cepora nerissa dapha ; Nakamura & Wakahara,2012 : 42.
Cepora nerissa dapha ; Onodera,2022 : 15, pl.63, figs.♂(wsf),♂(wsf,Un),♀(wsf),♀(wsf,Un),♂(dsf),♂(dsf,Un),♀(dsf),♀(dsf,Un). (Vientiane: Vang Vieng; Savannakhet: Don Natad; Xieng Khouang: Phou Samsoum)

[ CAMBODIA ]
Huphina dapha ; Vitalis,1919 : 218. (Cambodge)
Cepora nerissa dapha ; Kosterin,2019b : 26. (Ratanakiri)
Cepora nerissa dapha ; Kosterin,2020 : 313. (Siem Reap; Preah Vihear)
Cepora nerissa dapha ; Chartier & Kosterin,2022 : 114. (Preah Vihear; Ratanakiri; Siem Reap)

[ VIETNAM ]
Huphina nerissa dapha ; Godfrey,1919 : 464. (S: Daban)
Huphina nerissa nerissa ; Moulton,1923 : 132. (Pulo Condore)
Huphina nerissa dapha ; Metaye,1957 : 85. ("N" / C / S)
Cepora nerissa nerissa ; Ikeda et al.,1998 : 23, figs.8:5,♀(wsf),6,♂(wsf),7,♂(Un),8,♂(dsf),9,♂(Un),10,♀(dsf),11:42(♂genitalia). (Cuc Phuong)
Cepora nerissa dapha ; Luong et al.,2004 : 2, 31, fig.♂. (Cuc Phuong)
Cepora nerissa dapha ; Miyazaki, Saito & Saito,2006 : 11, figs.Pi-10,♂,♀. (Bin Thuan: Dong Tien; Ho Chi Minh)
Cepora nerissa dapha ; Monastyrskii & Devyatkin,2015 : 16. (C / S)
Cepora nerissa dapha ; Saito et al.,2017 : 78. (S: Nui Cam; Nui Coto; Nui Ket)
Cepora nerissa nerissa ; Monastyrskii,2024 : 54-56, pl.14, figs.2a♂(as ssp.nerissa),2b♂(Un, as ssp.nerissa),2c♂(as ssp.nerissa). (N: Muong Nhe; Cuc Phuong; Pu Luong; Xuan Son / C: Pu Mat; Vu Quang; Huong Son; Phong Nha-Ke Bang; Phong Dien/Dak Rong; Cu Lao Cham; Ngoc Linh) (in part)
Cepora nerissa dapha ; Monastyrskii,2024 : 54-56. (C: Chu Yang Sin; Nui Chua; Cat Loc / S: Cat Tien)

[ THAILAND ]
Mae Hong Son : 4♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 10♂ 4♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Lampang : 2♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phrae : 8♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 3♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchabun : 3♂ 2♀ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chaiyaphum : 2♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 7♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Bangkok : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Prachuap Khiri Khan : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchaburi : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Satun (Koh Tarutao) : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Yala : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Vientiane : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Champasak : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ VIETNAM ]
Lam Dong : 5♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is common in fields, open land, wasteland and roadside around secondary forests at low to moderate elevations (Alt. 10 - 1100m), and can also be found in the city. Both sexes visit flowers, and males frequently come to the banks of streams and puddles to suck water.
This taxon has very high seasonal and individual variation. Further taxonomic study is needed, including the relationship with the nominotypical subspecies.

Moore, F.,1879 : A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim,
with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4): 821-858, pls. 51-53.

Talbot, G.,1939 : Fauna of British India, including Ceylon & Burma. Butterflies. 1. (Papilionidae & Pieridae) (ed.2).
London : xxxix + 600pp, 1 map, 3 pls, 184 figs.

Yata, O., 1981 : Pieridae, in: Tsukada (Ed.), Butterflies of the South East Asian Islands 2.
Plapac co., Ltd., Tokyo. pp.205-438, pls.1-84. (in Japanese)

Mell, R.E., 1943 : Inventur und ökologisches Material zu einer Biologie der südchinesischen Pieriden.
Beiträge zur Fauna sinica 21.
Zoologica Stuttgart 36: 88-127.

♂ (dsf), Chiang Mai, Thailand.	♂ (wsf), Chiang Mai, Thailand.
♂ (dsf), Chiang Mai, Thailand.	♂ (dsf), Chiang Mai, Thailand.

1996.10.31 - 2025.04.30.