Leptosia nina nina

Leptosia nina nina (Fabricius, 1793)


♂, Mae Hong Son, Thailand.		♀, Mae Hong Son, Thailand.

♂, Bangkok, Thailand.		♀, Ranong, Thailand.

♂, Yala (Than To dist.), Thailand.		♀, Yala (Than To dist.), Thailand.

♂, Chanthaburi, Thailand.		♂, Satun, Thailand.

ON : Papilio nina Fabricius, 1793
OD : Ent. syst. 3(1):194-195.
TL : "India orientali" [S.India].

Distribution : Sri Lanka, India to Myanmar,
Thailand, Laos, Vietnam, Hainan, S.China,
Langkawi, W.Malaysia (Kedah; Penang).

	● ssp.nina
	● transition zone of nina and malayana
	● ssp.malayana

Synonym

Papilio xiphia Fabricius, 1781 : Spec. Ins. 2:43. TL. "India orientali" [S.India]. (homonym)

Records taken from Literature

[ THAILAND ]
Leptosia xiphia xiphia ; Fruhstorfer,1910 : 121. (Siam)
Leptosia xiphia xiphia ; Godfrey,1916 : 113.
Leptosia xiphia xiphia ; Pendlebury,1923 : 25. (Nakhon Sri Thammarat: Tai Sai River)
Leptosia xiphia xiphia ; Godfrey,1927 : 120. (N)
Leptosia xiphia xiphia ; Godfrey,1930 : 220-221. (all over)
Leptosia xiphia xiphia ; Godfrey,1931 : 192. (Ko Tao)
Leptosia xiphia xiphia ; Godfrey,1932a : 264. (P)
Leptosia nina nina ; Pinratana,1983 : 10, pl.9, figs.15a♂,♂(Un). (Chanthaburi; Chaiyaphum; Kanchanaburi; Nakhon Nayok; Lop Buri; Nakhon Ratchasima; Nonthaburi; Petchaburi; Saraburi; Phang Naga; Nakhon Sri Thammarat; Songkhla)
Leptosia nina nina ; Yamaguchi & Aoki,2000 : 251-252, fig.46,♂. (Mukdahan: Phu Mu)
Leptosia nina nina ; Yamaguchi & Aoki,2002 : 18. (Mukdahan: Ban Lao Kram; Ban Gok Tum)
Leptosia nina nina ; Yamaguchi, Aoki & Akiyama,2005 : 46. (Mukdahan: Nam Tok Tad Ton; Ubon Ratchathani: Don Far Huan)
Leptosia nina nina ; Katsuyama et al.,2005 : 65, fig.48,♂. (Umphang: Tha Sai; Umphang)
Leptosia nina nina ; Ek-Amnuay,[2007] : 138, pl.41, figs.Pi15a&♀,♀(Un). (in part)
Leptosia nina malayana ; Ek-Amnuay,[2007] : 138, pl.41, figs.Pi15b♂[Un],♂(Un)[Up]. (Yala; Khao Phanom Bencha)
Leptosia nina nina ; Les Day,2011-2022 : Samui Butterflies, figs.♂,♂(Un). (Koh Samui) (accessed: 2022.04.16)
Leptosia nina nina ; Kimura et al.,2011 : 154, figs.♂,♂(Un),♀,♀(Un). (Sarika; Nakohn Nayok; Wan Nam Khiewo; Phu Khiewo; Kao Soi Dao; Kanchanaburi; Khlong Naka; Muang Shone; Haadjai; Satun)
Leptosia nina malayana ; Kimura et al.,2011 : 154, figs.♂,♂(Un),♀,♀(Un). (Than To WF.)
Leptosia nina nina ; Ek-Amnuay,2012 : 138, pl.41, figs.Pi15a♀,♀(Un). (in part)
Leptosia nina malayana ; Ek-Amnuay,2012 : 138, pl.41, figs.Pi15b♂,♀. (Yala; Khao Phanom Bencha)
Leptosia nina nina ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Chiang Mai; Sa Kaeo; Bangkok; Saraburi; Chanthaburi; Phetchaburi) (accessed: 2024.04.13)
Leptosia nina malayana ; Küppers & Janikorn,2017 : 191, figs.Abb.Pi-024,033-034,118-122. (Phang Nga)
Leptosia nina nina ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Leptosia nina nina ; Motono & Negishi,1989 : 48, pl.15, figs.1,♂,2,♂(Un). (Tha Ngone; Vientiane)
Leptosia nina nina ; Osada, Uémura & Uehara,1999 : 202, pl.31, figs.♂,♀. (Thakhek; Nhahin)
Leptosia nina nina ; Nimura & Wakahara,2005 : 39. (Namdhua; Thakhek)
Leptosia nina nina ; Onodera,2009b : 34-35. (Vientiane: Namsong Riverside)
Leptosia nina nina ; Nakamura & Wakahara,2012 : 42.
Leptosia nina nina ; Onodera,2022 : 14, pl.61, figs.♂,♂(Un). (Vientiane: Vang Vieng)

[ CAMBODIA ]
Leptosia nina nina ; Woodfield & Murton,2006 : figs,♂,♂(Un). (Botum Sakor NP.)
Leptosia nina ; Onodera,2007 : 21,29, fig.10,♂. (Rattanakiri: Ta Veaeng; Siem Reap: Angkor)
Leptosia nina ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)
Leptosia nina nina ; Kosterin,2019 : 91. (Kampot)
Leptosia nina nina ; Kosterin,2019b : 26. (Ratanakiri)
Leptosia nina nina ; Kosterin,2020 : 314. (Siem Reap; Preah Vihear)
Leptosia nina nina ; Chartier & Kosterin,2022 : 113. (Koh Kong; Kampot; Phnom Penh; Preah Sihanouk; Pursat; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Leptosia xiphia xiphia ; Fruhstorfer,1910 : 121. (Annam)
Leptosia xiphia xiphia ; Godfrey,1919 : 464. (S: Daban)
Leptosia xiphia xiphia ; Moulton,1923 : 132. (Pulo Condore)
Leptosia xiphia ; Lemée,1950 : 63. (Saigon)
Leptosia xiphia xiphia ; Metaye,1957 : 87. (N / C / S)
Leptosia nina nina ; Ikeda et al.,1998 : 19, figs.6:19,♂,10:33(♂genitalia). (Cuc Phuong)
Leptosia nina ; Vu,2003 : 56. (Phu Quoc)
Leptosia nina ; Luong et al.,2004 : 2, 30, figs.♂,♂(Un). (Cuc Phuong)
Leptosia nina ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Leptosia nina nina ; Miyazaki, Saito & Saito,2006 : 11, figs.Pi-6,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien; Ta Cu; Ho Chi Minh)
Leptosia nina nina ; Nihira,2009 : 98. (Phu Quoc)
Leptosia nina nina ; Monastyrskii & Devyatkin,2015 : 16. (N / C / S)
Leptosia nina nina ; Saito et al.,2017 : 78. (S: Nui Cam; Nui Coto; Nui Ket)

Specimens Examined

[ THAILAND ]
Mae Hong Son : 3♂ 1♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chiang Rai : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chiang Mai : 12♂ (incl. photo & photo by M.Sugiyama).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Lampang : 5♂ (incl. photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phrae : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Nan : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Loei : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Tak : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Kanchanaburi : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phetchabun : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chaiyaphum : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Nakhon Nayok : 2♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Sa Kaeo : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Bangkok : 3♂ 2♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chanthaburi : 3♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Trat : 4♂ 1♀ (photo and photo by E.Ikari).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phetchaburi : 2♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Ranong : 9♂ 3♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Surat Thani (Koh Phangan Dist.) : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Krabi : 2♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Satun (Koh Tarutao) : 4♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Yala (Than To Dist.) : 20♂ 2♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ LAOS ]
Pakse : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ VIETNAM ]
Lam Dong : 2♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Notes

The butterfly is very common everywhere at low to high elevations (Alt. 10 - 1800m). Both sexes visit flowers and males occasionally come to the banks of streams and puddles to suck water.
The specimens from Than To, Yala have been considered as belonging to ssp.malayana, however after comparing a large number of specimens, they should be included in the nominotypical subspecies. The population is generally ssp.nina, however, individuals similar to ssp.malayana also occur rarely. It seems to be a transition zone of both subspecies.

Literature

Fabricius, J.C.,1781 : Species Insectorum exhibentes eorum differentias specificas, synonyma auctorum, locanatalia, metamorphosin adjectis observationibus descriptionibus.
vol.2:1-494. Humburg and Cologne, Karl E.Bohn.

Fabricius, J.C.,1793 : Entomologia systematica emedata et aucta. secundum classes, ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus.
3(1): [iv] + 488 pp. Copenhagen. C.G.Proft.

Corbet, A.S.,1941 : Observations on Certain of the Fabrician Names of Indo-Australian Rhopalocera (Lepid.)
Proc. R. ent. Soc. Lond. (B)10(6):98-106.


♂, Chiang Mai, Thailand. (photo by M.Sugiyama)	♂, Chiang Mai, Thailand.

♂+♀ (mating), Trat, Thailand.

1996.10.31 - 2024.04.30.