Graphium antiphates alcibiades

Graphium antiphates alcibiades (Fabricius, 1787)


♂, Yala, Thailand.		♀, Trang, Thailand. (photo by A.M.Cotton)

♂, Phuket, Thailand. (photo by A.M.Cotton)		♂, Narathiwat, Thailand. (photo by A.M.Cotton)	♂, Narathiwat, Thailand. (photo by A.M.Cotton)

♂, Ranong, Thailand. (photo by A.M.Cotton)	♂, Ranong, Thailand. (photo by A.M.Cotton)	♂ genitalia, Narathiwat, Thailand.


♂, Phetchaburi, Thailand. (photo by A.M.Cotton)	♂, Phetchaburi, Thailand. (photo by A.M.Cotton)	♂, Phetchaburi, Thailand. (photo by A.M.Cotton)	♂, Phetchaburi, Thailand. (photo by A.M.Cotton)

ON : Papilio alcibiades Fabricius,1787
OD : Mantissa Ins. 2: 8.
TL : 'Tranquebariae' [loc. err. = Malay Peninsula]. (ZMUC)

Distribution : P.Thailand (South from Phetchaburi), W.Malaysia, Singapore, Bintan.

	● ssp.nebulosus
	● ssp.antiphates
	● ssp.matsui
	● transition zone of nebulosus and alcibiades
	● ssp.alcibiades
	● ssp.toshikazui

Synonym

Arisbe itamputi tungi Page,2020 : Nachr. ent. Ver. Apollo, N.F. 40(3/4): 194-196, figs.5e(HT,♂), 5f(PT), 5g(PT). TL. Taiping, Perak, Malaysia. (ETHZ)

Type Material Information

alcibiades ; Syntypes, 2♂. (ZMUC, photo examined.)
tungi ;
Holotype, ♂, 3.Apr.1976. Sengei Batu Tegeh, Taiping, Perak, Malaysia. (ETHZ)
Paratypes : (36♂ 1♀)
Thailand : 3♂, Ranong; 1♂, Ko Samui; 1♂, Nakhon si Thammarat; 6♂, Phuket; 3♂, Trang;
W.Malaysia : 5♂, Sengei Batu Tegeh, Taiping, Perak; 2♂, Ipoh, Perak; 6♂ 1♀, Jelebu, Negri Sembilan; 1♂, Tapah Hills, Perak; 2♂, Jerantut, Pahang; 4♂, Taman Negara Rompin, Johor;
Singapore : 1♂ ; Indonesia: 1♂, Bintan.

Records taken from Literature

[ THAILAND ]
Graphium antiphates itamputi ; Pinratana,1977 : 54, figs.P41b♂,♂(Un). (P)
Graphium antiphates itamputi ; Pinratana & Eliot,1992 : 33,45, pl.58, fig.48b♂. (Ranong; Trang; Yala)
Graphium antiphates pompilius ; Nishimura,1996a : 45-47, figs.23,♂,24,♂. (Nakhon Si Thammarat: Namtok Ka Rom; Phuket: Namtok Tone Sai; Songkhla: Namtok Ton Nga Chang; Yala: Than To)
Graphium antiphates itamputi ; RDPB & DNP,2006-2008 : Butterflies in Hala-Bala Forest, fig.[♂](Un). (Narathiwat: Hala Bala) (last update: 2006.12.31 ; accessed: 2024.04.10)
Pathysa antiphates itamputi ; Ek-Amnuay,[2007] : 112, pl.28, figs.P56b♂,♂(Un). (Ranong)
Graphium antiphates itamputi ; Kimura et al.,2011 : 133, figs.♂,♂(Un). (Khlong Naka; Kam Puan; Ka Chong; Than To WF.) (in part)
Graphium antiphates pompilius ; Ek-Amnuay,2012 : 112, pl.28, figs.P53,♂(Lower row),♂(Lower row, Un). (Ranong) (in part)
Arisbe antiphates pompilius ; Küppers & Janikorn,2017 : 169-170, figs.Abb.Pa-140-141, 290-306. (Phang Nga)
Arisbe itamputi tungi ; Page,2020 : 194-196. (Ranong; Nakhon si Thammarat; Phuket; Trang) (in part)
Graphium antiphates alcibiades ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2022- : Butterflies of Thailand, fig. (Yala) (accessed: 2023.05.18)

Specimens Examined

[ THAILAND ]
Phetchaburi : 12♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Ranong : 13♂ 2♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phuket : 2♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Trang : 1♂ 1♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Pattani : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Satun : 2♂ (photo by E.Ikari and A.Giudici).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Yala : 12♂ 1♀ (incl. photo & photo by S.Atdhabhan).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Narathiwat : 6♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Notes

The butterfly is common in montane forests at low to moderate elevations (Alt. 10 - 800m). Males frequently come to the banks of streams and puddles to suck water.
Page (2020) separated itamputi from antiphates based on DNA analysis (Makita et al.,2003, Wilson et al.,2014). Furthermore, he treated alcibiades as the continental subspecies of antiphates, and described Arisbe itamputi tungi for the populations from P. Thailand to Singapore and Bintan.
However, both Makita et al. (2003) and Wilson et al. (2014) made no mention or conclusion about the relationship between antiphates and itamputi. We cannot find any notable differences in the genitalia of each taxon, and there is a transition zone of both taxon in northern part of Peninsular Thailand. Thus, here we provisionally follow the traditional classification. Further taxonomic study is needed.
Cotton (2018) presumed the type locality of alcibiades is somewhere in Malay Peninsula, possibly Phuket, P. Thailand. I agree with his opinion and consider tungi to be a synonym of alcibiades.
The population from Phetchaburi, northern part of Peninsular Thailand is rather variable. Generally ssp.alcibiades, but intermediate individuals between ssp.nebulosus are often found. It seems to be a transition zone between both subspecies.

Literature

Fabricius, J.C.,1787 : Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis,
differentiis specificis, emendationibus, observationibus.
Copenhagen, Christ. Gottl. Proft 8vo 1: xx+348pp ; 2: [2]+382pp

Butler, A.G.,1885 : [ Description of Papilio itamputi ].
In H.O.Forbes, A Naturalist's Wanderings in the Eastern Archipelago, a narrative of Travel and exploration from 1878-1883.
London : xix + 536pp, 22 pls, 4maps (8vo).

Fruhstorfer, H.,1909 : Neue Papilioniden des indo-australischen Faunengebietes.
Ent. Zeit. 22(41): 169-170. (9.Jan.1909)

Jordan, K.,1909 : Die Indo-Australische Tagfalter. Papilionidae in Seitz, Grossschmett. Erde
9: 89-96.

Nishimura, M.,1996 : Notes on some Papilionidae from Indo-China. (2).
Tyô to Ga. 47(1): 40-48. (in Japanese)

Makita, H., Shinkawa, T., Kondo, K., Xing, L. & Nakazawa, T.,2003 : Phylogeny of the Graphium butterflies inferred from nuclear 28S rDNA and mitochondrial ND5 gene sequences.
Trans. lepid. Soc. Jap. 54(2): 91–110.

Wilson, J.J., Karen-Chia, H.M., Sing, K. & Azirun, M.,2014 : Towards resolving the identities of the Graphium butterflies (Lepidoptera: Papilionidae) of Peninsular Malaysia.
J. Asia Pac. Entomol. 17: 333–338.

Cotton, A.M.,2018 : Confirmation of the true identity of Papilio alcibiades Farbicius, 1787 (Lepidoptera: Papilionidae).
Butterflies 79: 11-17.

Page, M.G.P.,2020 : Notes on Arisbe (Pathysa) antiphates (Cramer, 1775) (Lepidoptera, Papilionidae) and its allies.
Nachr. ent. Ver. Apollo, N.F. 40(3/4): 187-202.


♂, Yala, Thailand. (photo by S.Atdhabhan)	♂, Satun, Thailand. (photo by A.Giudici)

♂, Yala, Thailand.

2014.07.19 - 2024.04.30.