Atrophaneura varuna zaleucus   (Hewitson,[1865])

Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♂ Un. Atrophaneura varuna zaleucus ♀ Up. Atrophaneura varuna zaleucus ♀ Un.
♂, 1988.02.15. Wang Chin, Phrae, Thailand. ♀, 1993.09.24. Doi Phu Kha, Nan, Thailand.
Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♀ Up.
♂, 1989.09.--. Doi Phu Kha, Nan, Thailand. ♂, 1989.02.01. Chiang Dao, Chiang Mai, Thailand. ♂, 1990.01.--. Chiang Dao, Chiang Mai, Thailand. ♀, 2006.04.27. Ban Koun Ngun, Hinboun, Laos.
( photo by A.M.Cotton )
Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♂ Un. Atrophaneura varuna zaleucus ♂ Up. Atrophaneura varuna zaleucus ♂ Un.
♂, 2006.05.09. Nong Ya Plong, Phetchaburi, Thailand.
( photo by A.M.Cotton )
♂, 2007.08.26. Nong Ya Plong, Phetchaburi, Thailand.
( photo by A.M.Cotton )

transition form ssp.zaleucus to ssp.astorion
Atrophaneura varuna zaleucus ♀ Up. Atrophaneura varuna zaleucus ♀ Un. Atrophaneura varuna zaleucus ♀ Up. Atrophaneura varuna zaleucus ♀ Un.
♀, 1992.06.12. Padeng, Phetchaburi, Thailand. ♀, 2004.08.1. Dalat, Lam Dong, Vietnam.( photo by K.Saito )
Atrophaneura varuna zaleucus ♀ Up. Atrophaneura varuna zaleucus ♀ Un.
♀, 1990.05.20. Wiang Pa Pao, Chiang Rai, Thailand.
( photo by A.M.Cotton )

ON : Papilio zaleucus  Hewitson,[1865]
OD : Ill. exot. Butts. 3:[5], pl.[3], figs.24(♂),25(♀) (Papilio VIII).
TL : Burmah. (BMNH)
Distribution : Myanmar, Thailand, Laos, Cambodia, Vietnam, China.
Atrophaneura varuna zaleucus ♂ genitalia Atrophaneura varuna zaleucus map

Synonyms

* Byasa zaleucus ♂v.punctata  Evans,1923 : J. Bombay nat. Hist. Soc. 29(1):231.  TL. Lower Burma. (BMNH)
Papilio zaleucus f.indiv.♂anomala  Rousseau-Decelle,1947 : Bull. Soc. ent. Fr. 51:128.  TL. "Kalataung, Tavoy, Birmanie" (♂)
Atrophaneura varuna liziensis  Zhao,1997 : in Chao & Wang, Lepidoptera of China 3 : fig.♂,♂(Un).  TL. Lizi, Xizhang.


Type Materials Information

zaleucus ;
♂, Type (red), Rh10831, Burmah. (BMNH. examined.) ; ♀, Type (red), Rh10832, Burmah. (BMNH. examined.)
punctata ; ♂, Type (red), Rh10830, Kawkareik, Thinyanyaung, Tenasserim. 1922.Dec. (BMNH. examined.)
liziensis ; ♂, Holotype, 1992.Vii., Lizi, Xizhang. (Chengdu Museum)


Records taken from Literature

[ THAILAND ]
Papilio zaleucus ; Godfrey,1916 : 109. (Chonburi: Hup Bon)
Papilio zaleucus ; Godfrey,1927 : 119. (North)
Papilio zaleucus ; Godfrey,1930 : 207. (Prae: Me Song forest / Chiang Mai / Chonburi: Sriracha / West: Me Wong)
Papilio zaleucus ; Reeves,1967a : 120. (Nakhon Nayok: Khao Yai)
Atrophaneura varuna zaleucus ; Pinratana,1977 : 11, fig.P6c,♂,♀. (exept Peninsula)
Parides varuna zaleucus ; Pinratana,1992 : 13,39, pl.9, fig.6,♂,♀. (Chiang Mai / Chaiyaphum / Chonburi / Chanthaburi / Ratchaburi)
Atrophaneura zaleucus ; Katsuyama et al.,2005 : 64, fig.37♂. (Umphang: Tha Sai ; Umphang)
Atrophaneura zaleucus ; Katsuyama et al.,2006 : 45, fig.23♂. (Nan: Ban Nam Poon)
Atrophaneura zaleucus ; Ek-Amnuay,[2007] : 66, pl.4, fig.P8,♂,♂(Un),♂,♀,♀(Un). (Chiang Mai / Chiang Rai)
Atrophaneura varuna zaleucus ; Racheli & Cotton,2010 : 29-31. (continental Thailand)
Atrophaneura varuna astorion ; Kimura et al.,2011 : 114, fig.♀,♀(Un). (Ratchaburi)
Atrophaneura zaleucus ; Kimura et al.,2011 : 113, fig.♂,♂(Un),♂,♂(Un), ♀,♀(Un). (Khun Lao Village / Maethro / Bang Salok / Thung Saleng Luang / Phu Khiewo / Nakohn Nayork / Wan Nam Khiewo / Kao Soi Dao / Pliewn WF / Khao Sabap / Kratchin WF / Khao Khiewo / Sai York Noi / Ratchaburi)
Atrophaneura varuna zaleucus ; Ek-Amnuay,2012 : 66, pl.4, fig.P8a,♂,♂(Un),♂,♀,♀(Un). (Chiang Mai / Chiang Rai / Kaeng Krachan / Khao Ang Rue Nai)

[ LAOS ]
Papilio zaleucus ; Dubois & Vitalis de Salvaza,1914 : 146. (Laos)
Papilio zaleucus ; Dubois & Vitalis de Salvaza,1921 : 8. (Laos)
Atrophaneura varuna zaleucus ; Motono & Negishi,1989 : 45, not fig. (Nam Tok Tat / Luang Prabang / Vang Vieng)
Atrophaneura zaleucus ; Masui & Uehara,1994 : 8, pl.2, figs.1♂,2♀. (Louang Namtha)
Atrophaneura zaleucus ; Osada, Uémura & Uehara,1999 : 200, pl.3, fig.♂. (Vang Vieng)
Atrophaneura varuna zaleucus ; Cotton & Racheli,[2007] : 326-327. (Thabok / Phou Pha Man: Koun Ngun)
Atrophaneura varuna zaleucus ; Racheli & Cotton,2010 : 29-31, fig.♀,♀,♀(Un). (W.Laos)
Atrophaneura zaleucus ; Nakamura & Wakahara,2012 ; 41.

[ CAMBODIA ]
Atrophaneura varuna zaleucus ; Racheli & Cotton,2010 : 29-31. (W.Cambodia)

[ VIETNAM ]
Papilio zaleucus ; Jordan,1909 : 30. (Tonkin)
Papilio zaleucus ; Dubois & Vitalis de Salvaza,1914 : 146. (Annam)
Papilio zaleucus ; Dubois & Vitalis de Salvaza,1921 : 8. (Tonkin / Annam)
Papilio zaleucus ; Metaye,1957 : 81. (North / Central)
Atrophaneura zaleucus ; Monastyrskii,2007 : 27, pl.3, figs.1a♂,1b♀. (North: Muong Nhe / Central: Cua Rao (Annam) / South: Huong-Va)
Atrophaneura varuna zaleucus ; Racheli & Cotton,2010 : 29-31. (NW.Vietnam)


Specimens Examined

[ THAILAND ]
Mae Hong Son, Ban Ten Doi.
1988.03.24


1♀
Y.Inayoshi (YI)
Chiang Rai, Mae Kajan.
1987.02.--
2♂
1♀
Local Catcher (YI)
Chiang Rai, Mae Kajan.
1987.03.--
1♂


Local Catcher (YI)
Chiang Rai, Mae Kajan.
1987.04.01
3♂


Local Catcher (YI)
Chiang Rai, Wiang Pa Pao.
1990.05.20


1♀
Local Catcher (AC)
Chiang Mai, Chiang Dao.
1990.01.--
1♂
1♀
Local Catcher (MN)
Chiang Mai, Chiang Dao.
1989.02.01
1♂
1♀
ocal Catcher (MN)
Chiang Mai, Chiang Dao.
1977.03.07
1♂


P.Sukkit (PS)
Chiang Mai, Chiang Dao.
1977.03.15
1♂


P.Sukkit (PS)
Chiang Mai, Chiang Dao.
1979.05.02


1♀
P.Sukkit (PS)
Chiang Mai, Doi Suthep.
1983.04.04


1♀
Y.Inayoshi (YI)
Chiang Mai, Chiang Mai dist.
2013.08.17


1♀
E.Ikari (EI)
Phrae, Wang Chin.
1988.02.15
1♂


Local Catcher (YI)
Phrae, Wang Chin.
2007.03.01


1♀
Local Catcher (AC)
Phrae, Wang Chin.
2012.10.26
1♂


Y.Inayoshi (YI)
Lampang, Jae Hom.
1988.02.02
1♂


Local Catcher (YI)
Lampang, Jae Hom.
1988.02.26
1♂


Local Catcher (YI)
Nan, Doi Phu Kha.
1991.02.--
1♂


Local Catcher (YI)
Nan, Doi Phu Kha.
1994.02.22


1♀
P.Sukkit (PS)
Nan, Doi Phu Kha.
1993.03.04
2♂
1♀
P.Sukkit (PS)
Nan, Doi Phu Kha.
1991.03.12
1♂


Y.Inayoshi (YI)
Nan, Doi Phu Kha.
1991.03.12
1♂


Y.Inayoshi (YI)
Nan, Doi Phu Kha.
1991.03.13
1♂


Y.Inayoshi (YI)
Nan, Doi Phu Kha.
1994.08.14


1♀
P.Sukkit (PS)
Nan, Doi Phu Kha.
1990.09.--
1♂


P.Sukkit (PS)
Nan, Doi Phu Kha.
1993.09.24


1♀
Local Catcher (YI)
Nan, Doi Phu Kha.
1993.10.11


1♀
Local Catcher (YI)
Chonburi, Khao Khiew.
1976.06.06


1♀
P.Sukkit (PS)
Chanthaburi, Khao Soi Dao.
1983.03.27
2♂


Y.Inayoshi (YI)
Chanthaburi, Khao Soi Dao.
1983.03.28
1♂


Y.Inayoshi (YI)
Chanthaburi, Khao Soi Dao.
1984.12.20
4♂


P.Sukkit (PS)
Chanthaburi, Khao Soi Dao.
1984.12.21
1♂


P.Sukkit (PS)
Phetchaburi, Padeng.
1992.06.12


1♀
Local Catcher (MN)
Phetchaburi, Nong Ya Plong.
2006.05.09
1♂


A.Rukkasikorn (AC)
Phetchaburi, Nong Ya Plong.
2007.08.26
1♂


A.Rukkasikorn (AC)

[ LAOS ]
Thabok, Tad Xai WF.
2006.02.21
1♂


A.M.Cotton (AC)
Hinboun, Ban Koun Ngun.
2006.04.27


1♀
A.M.Cotton (AC)

[ VIETNAM ]
Lam Dong, Dalat.
2004.08.15


1♀
K.Saito (KS)


Notes

Figured specimen from Padeng, Phetchaburi seems a transition form ssp.astorion to ssp.zaleucus.
In the BMNH, There are several specimens (from S.Myanmar : Tavoy and Tenasserim.) same as this form.


Literature

Hewitson, W.C.,[1862]-[1866] :
Illustrations of new species of exotic Butterflies, selected chiefly from the Collections of W.Wilson Saunders and William C. Hewitson.
3:iv + [124]pp, 60 pls. Van Voorst, London.

Rousseau-Decelle, G.,1947 : Contribution a letude des Papilio de la faune Indo-Oceanienne (Lep. Papilionidae)
Bull. Soc. ent. Fr. 51:128-133.

Zhao, L.,1997 : Atrophaneura varuna liziensis Zhao (subspecies nov.),
in Chao, L. & H.Y.WANG, Lepidoptera of China 3. Papilionidae, Danaidae, Pieridae, Amathusiidae.
The Taiwan Museum, Taipei.

1996.09.26 - 2014.05.31.