Limenitis dudu secius
(Tsukada, 1991)
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♂, Chiang Mai, Thailand. |
♀, Chiang Mai, Thailand. |
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♂, Nan, Thailand. |
♀, Nan, Thailand. |
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♂, Loei, Thailand. |
♀, Nakhon Nayok, Thailand. |
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♂, Vinh Phuc, Vietnam. |
♀, Vinh Phuc, Vietnam. |
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♂, Nan, Thailand. |
♂, Gia Lai, Vietnam. (photo by K.Saito) |
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♂, Houaphan, Loas. (photo by K.Saito) |
♂, Lam Dong, Vietnam. (photo by K.Saito) |
ON :
Sumalia dudu secius Tsukada, 1991
OD : In: Tsukada, Butterflies of the South East Asian Islands. 5: 297,543, pl.18, figs.4,(PT,♂Un), 7(HT,♂).
TL : "Tamdao, N.Vietnam" [Tamdao, Vinh Phuc, N.Vietnam]. (in coll. E.Tsukada)
Distribution : Thailand, Laos, Vietnam, S.China.
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Synonym
Sumalia dudu pilosus Tsukada, 1991 : In: Tsukada, Butterflies of the South East Asian Islands. 5: 297,543, pl.18, figs.1(HT,♂), 2(PT,♂Un), 3(PT,♀). TL. "Chieng Dao, N.Thailand" [Chiang Dao, Chiang Mai, N.Thailand]. (in coll. E.Tsukada)
Type Materials Information
secius :
Holotype ♂, 11.Apr.1990. Tamdao, 1,300m, N.Vietnam. (in coll. E.Tsukada)
Paratypes ; 3♂, same locality as holotype.
philosus :
Holotype ♂, Feb.1980. "Chieng Dao" [Chiang Dao, Chiang Mai], N.Thailand. (in coll. E.Tsukada)
Paratypes ; all paratypes from Thailand.
4♂, "Chieng Dao" [Chiang Dao, Chiang Mai] ; 4♂ 1♀, "Phu Khies" [Phu Khieo, Chaiyaphum] ; 1♂, Nan ; 1♂, Fang, [Chiang Mai] ; 4♂, "Vieng Pa Pao" [Wiang Pa Pao, Chiang Rai].
Records taken from Literature
[ THAILAND ]
Limenitis dudu ; Godfrey,1932b : 325. (Chiang Mai: Doi Suthep)
Sumalia dudu dudu ; Pinratana,1979 : 62, figs.N110,♂,♂(Un). (Chiang Mai; Kao Yai)
Sumalia dudu pilosus ; Tsukada,1991 :297,543, pl.18, figs.1(HT,♂),2(PT,♂Un),3(PT,♀). ("Chieng Dao" [Chiang Dao]; "Phu Khies" [Phu Khieo]; Nan; Fang; "Vieng Pa Pao" [Wiang Pa Pao])
Limenitis dudu dudu ; Pinratana & Eliot,1996 : 63, pl.41, fig.115,♂. (Chiang Mai; Phisanulok; Nakhon Nayok)
Parasarpa dudu dudu ; Ek-Amnuay,[2007] : 412, pl.180, figs.N120,♂,♂(Un),♀. (Doi Pha Hom Pok)
Parasarpa dudu dudu ; Ek-Amnuay,2012 : 422, pl.184, figs.N286,♂,♂(Un),♀. (Doi Pha Hom Pok; Doi Phu Kha)
Parasarpa dudu dudu ; Kimura et al.,2016 : 93, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Pha Hom Pok; Doi Phui; Doi Suthep; Doi Inthanon; Wang Chin; Thung Saleng Luang)
Limenitis dudu secius ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2024- : Butterflies of Thailand, figs., Early Stages. (Chiang Mai; Kamphaeng Phet; Phetcahabun; Nakhon Nayok) (accessed: 2024.05.13)
[ LAOS ]
Sumalia dudu dudu ; Masui & Uehara,1995 : 5, pl.2, fig.2,♂. (Phong Sali)
Parasarpa dudu dudu ; Osada, Uémura & Uehara,1999 : 207, pl.61, fig.♂. (Oudomxay)
Parasarpa dudu dudu ; Nakamura & Wakahara,2012 : 46, pl.4, figs.Ny-103,♀,♀(Un). (Phou Samsoum)
Parasarpa dudu dudu ; Onodera,2022 : 40, pl.119, figs.♂,♂(Un). (Bolikhamxai: Thabok)
[ VIETNAM ]
Limenitis dudu f. dudu ; Dubois & Vitalis,1924 : 41. (Tonkin: Chapa; Cochinchine: Gyarai)
Limenitis dudu dudu ; Metaye,1957 : 97. (N / S)
Parasarpa dudu ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Bach Ma)
Parasarpa dudu hainensis ; Miyazaki, Saito & Saito,2006b : 26, figs.N-75,♂,♀. (Lam Dong: Dambri)
Parasarpa dudu dudu ; Yokochi et al.,2015 : 41. (Khanh Hoa: Hon Ba)
Parasarpa dudu dudu ; Monastyrskii & Devyatkin,2015 : 36. (N / C: to Lam Dong)
Parasarpa dudu dudu ; Monastyrskii,2019 : 4(2):41-42, pl.40, figs.1a♂,1b♂(Un). (N: Hoang Lien; Van Ban; Dong Van; Dong Phuc; Lung Li; Ba Vi; Tam Dao / C: Bach Ma; Son Thanh; Bi Doup-Nui Ba; Hon Ba; Chu Yang Sin; Loc Bac)
Specimens Examined
[ THAILAND ]
Chiang Mai : 14♂ 2♀ (incl. photo).
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Nan : 15♂ 3♀.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Loei : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Chaiyaphum : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Nakhon Nayok : 1♀ (photo by S.Sophonviwatkul).
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
[ LAOS ]
Houaphan : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
[ VIETNAM ]
Vinh Phuc : 2♂ 1♀.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Gia Lai : 1♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Lam Dong : 3♂.
| Jan | Feb | Mar | Apr | May | Jun |
Jul | Aug | Sep | Oct | Nov | Dec |
Notes
The butterfly is locally common in montane forests at moderate to high elevations (Alt. 900 - 1800m).
Tsukada (1991) described ssp.philosus from Thailand and ssp.secius from Vietnam.
Certainly N. Thailand specimens tend to have a wider band and the Vietnamese specimens have a narrower band.
However, the individuals from Thailand, Laos and Vietnam show more seasonal and individual variation than geographical variation, so it is not possible to make a clear distinction.
Therefore, I consider all Indochinese populations to be secius and treat philosus as a synonym.
Literature
Tsukada, E.,1991 : In: Tsukada, E. (Ed.), Butterflies of the South East Asian Islands 5. Nymphalidae (II).
Nagano-ken, Azumino Butterflie's Research Institute. 576 pp. (238 col. pls.) (in Japanese)
Wu, L.W., Chiba, H., Lees, D.C., Ohsima, Y. & Jeng, M.L.,2019 : Unravelling relationships among the shared stripes of sailors:
Mitogenomic phylogeny of Limenitidini butterflies (Lepidoptera, Nymphalidae,Limenitidinae), focusing on the genera Athyma and Limenitis.
Mol. Phylogenet. Evol. 130(2019): 60-66.
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♂, Chiang Mai, Thailand. |
♂, Chiang Mai, Thailand. |
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♂, Chiang Mai, Thailand. |
♂, Loei, Thailand. |
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♂, Nakhon Nayok, Thailand. (photo by T.Aoyama) |
♂, Chiang Mai, Thailand. |
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♀, ♀(laying egg), Nakhon Nayok, Thailand. (photo by T.Aoyama) |
A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.
