♂, Lam Dong, Vietnam. (photo by K.Saito)		♀, Bolikhamxai, Laos. (photo by H.Onodera)
♂, Bolikhamxai, Laos. (photo by H.Onodera)

ON : Isma bononia ssp. OD : TL : Distribution : S.E.Thailand, Laos, Vietnam. ● ssp. ● ssp.bononia

[ THAILAND ]
Sepa inarime ; Godfrey,1930 : 376. (Chonburi: Hup Bon)
Isma bononia idyalis ; Kimura et al.,2011 : 77, figs.♂,♂(Un). (Kao Soi Dao)

[ LAOS ]
Isma bononia ssp. ; Masui & Uehara,1995 ; 5, pl.2, fig.13,♂,♂(Un). (Saravan)
Isma bononia ssp. ; Osada, Uémura & Uehara,1999 : 224, pl.140, figs.♂,♀. (Phong Si)
Isma bononia idyalis ; Nakamura & Wakahara,2012 : 59.
Isma bononia idyalis ; Onodera,2022 : 118, pl.253, figs.♂,♂(Un),♀,♀(Un). (Khammouan: Nahin)

[ VIETNAM ]
Isma bononia idyalis ; Devyatkin & Monastyrskii,2002 : 149. (Ba Be; Huu Lien; Cuc Phuong; Ke Bang; Kon Cha Rang)
Isma bononia idyalis ; Devyatkin & Monastyrskii,2003 : 125. (S: Cat Tien)
Isma bononia ssp. ; Miyazaki, Saito & Saito,2007b : 14, fig.H-83,♂. (Lam Dong: Dambri; Dai Duc Me)
Isma bononia bononia ; Miyazaki, Saito & Saito,2007c : 40, fig.H-83,♀. (Lam Dong: Dambri)
Isma bononia idyalis ; Monastyrskii & Devyatkin,2015 : 80. (N / C: to Lam Dong)

[ Laos ]
Bolikhamxai : 1♂ 1♀ (photo by H.Onodera).

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[ VIETNAM ]
Lam Dong : 1♂.

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The populations from S.E.Thailand, Laos and Vietnam have often been treated as ssp.idyalis. However, the males have a conspicuous white spot on the hindwing, a feature that is not present in ssp.idyalis. It is also unlikely that they are the same taxon based on biogeography. Further taxonomic study is needed.

Nicéville, L.de,1897 : On new or little-known Butterflies from the Indo- and Austro-Malayan Regions.
J. asiat. Soc. Bengal, pt 2, 66(3):543-577, pl.1-4.

2017.12.31 - 2023.08.31.