Isma guttulifera kuala  (Evans,1932)

Isma guttulifera kuala ♀ Up. Isma guttulifera kuala ♀ Un.
♀, Ranong, Thailand.

ON : Sepa guttulifera kuala Evans,1932
OD : Ident. Indian Butts. (2nd ed.) : 379.
TL : Pahang, Malaysia. (♂, BMNH)
Distribution : P.Thailand, W.Malaysia, Borneo.

Records taken from Literature

Isma kuala ; Pinratana,1985 : 76, pl.26, fig.162. (Yala)
Isma guttulifera kuala ; Ek-Amnuay,[2007] : 798. (Trang)
Isma guttulifera ; Ek-Amnuay et al.,2007 : 5, fig.H189,♀ (in part). (Ranong: Kam Puan)
Isma guttulifera ; Kimura et al.,2011 : 77. (by literature)
Isma damocles ; Kimura et al.,2011 : 77, fig.♀,♀(Un) (in part). (Kam Puan)
Isma guttulifera ; Ek-Amnuay,2012 : 854, pl.394, fig.H186,♀ (in part).

Specimens Examined

Ranong : Suk Samran dist. 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Identification about female of I.damocles and I.guttulifera is still not clear.
The difference between these species, the upper side of the forewing, spot in space 2 overlapped to lower cell spot or not.
Although Eliot,1992 treated overlapped species as guttulifera and another species as damocles, Maruyama,1991 treated them exactly the opposite.
For the time being, I follow the treatment of Maruyama.


Evans, W.H.,1932 : The identification of Indian butterflies. second edition revised.
Bombay nat. Hist. Soc., Madras : 10+454pp, 32 pls, 9f.

Maruyama, K.,1991 : Butterflies of Borneo vol.2, No.2. Hesperiidae, Addndum of vol.1.
XII+89 pp(Japanese text), 48 pls, XII+83 pp (English text) + 1.
Tobishima corporation. Tokyo, Japan.

1999.09.11 - 2017.12.31.