♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Chanthaburi, Thailand.		♀, Chanthaburi, Thailand.
♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Lao Cai, Vietnam. (photo by K.Saito)		♀, Lao Cai, Vietnam. (photo by K.Saito)
♂, Chiang Mai, Thailand.		♂, Kien Giang, Vietnam. (photo by K.Saito)
♂ (f. layardi), Kien Giang, Vietnam. (photo by K.Saito)		♂ (f. layardi), Kien Giang, Vietnam. (photo by K.Saito)
♂ genitalia, Vientiane, Laos.

ON : Euploea godartii Lucas, 1853 OD : Revue Mag. Zool. (2)5:319-321. TL : ['Java'] Distribution : Myanmar, Thailand, Laos, Cambodia, Vietnam. ● ssp.godartii ● transition zone of godartii and amymone ● transition zone of godartii and graminifera ● ssp.graminifera

Euploea siamensis C. Felder & R. Felder, 1865 : Reise öst. Fregatte Novara. Zool. Theil (2):341-342, pl.41, fig.6,♂. TL. Siam ; Cochin.
Euploea layardi H.Druce, 1874 : Proc. zool. Soc. Lond. 1874(1):103-104, pl.16, fig.1,♀. TL. Chentaboon [Chanthaburi], Siam. (NHML)
Euploea subdita Moore, 1879 : Proc. zool. Soc. Lond. 1878(4):823. TL. Hatsiega, Tenasserim. (NHML)
Crastia binghami Moore, 1883 : Proc. zool. Soc. Lond. 1883(3):278-279. TL. Thoungyeen, Upper Tenasserim. (NHML)
Crastia circuita Swinhoe, 1903 : Ann. Mag. nat. Hist. (7)11:499. TL. Tonkin. (NHML)
Crastia tonkinensis Swinhoe, 1903 : Ann. Mag. nat. Hist. (7)11:499. TL. Tonkin. (NHML)
Euploea godarti f. dominans Fruhstorfer, 1910 : in Seitz, Grossschmett. Erde 9:236. TL. Siam. (NHML)
Euploea godarti f. recussa Fruhstorfer, 1910 : in Seitz, Grossschmett. Erde 9:236. TL. S.Vietnam. (NHML)
Euploea godarti f. defigurata Fruhstorfer, 1910 : in Seitz, Grossschmett. Erde 9:236. TL. U.Burma.
Euploea core f. penanga Talbot, 1940 : Ent. mon. Mag. 76:248. TL. Penang Hill, Malaya.
Euploea core garampiana Murayama, 1961 : Trans. lepid. Soc. Jap. 11(4):56, 59, fig.4,♀. TL. Garampi, Formosa. (Southern part of Formosa).

'godartii ; ♀, Possible Type HT (red), Ind. or. Ex. Typicalibus Speciminibus. Ex. Musaeo Boisduval (Euploea siamensis Felder (novara, 1, tab.XLI, no.6)' (NHML, examined)
layaardi ; ♀, Type (red), Rh6621, Chentaboon [Chanthaburi], Siam. (NHML, examined)
subdita ; ♂, Type (red), Rh6617, Upper Tenasserim. (NHML, examined)
binghami ; ♂, Type (red), Rh6618, Thoungyan Valley, Tenasserim. Capt.Bingham ; ♀, Type (red), Rh6619, Thoungyeen Forests, Burma. Capt. Bingham (NHML, examined)
circuita ; ♂, Type (red), Rh6622, Tonkin. E.Swinhoe (NHML, examined)
tonkinensis ; ♂, Type (red), Rh6624, Tonkin. E.Swinhoe (NHML, examined) ; ♀, Type (red), Rh6623, Tonkin. E.Swinhoe (NHML, examined)
dominans ; ♂, Type, Jan. Muok Lek, Siam. H.Fruhstorfer (NHML, examined)
garampiana ; ♀, Type, July.1959. Garampi (Southern part of Formosa)

[ THAILAND ]
Euploea godarti godarti [sic] ; Fruhstorfer,1910 : 236. (Ko Chang; Siam)
Euploea godarti [sic] tonkiniensis ; Fruhstorfer,1910 : 236. (Siam)
Euploea godarti [sic] ; Godfrey,1916 : 119-120.
Euploea orontobates ; Godfrey,1916 : 120. (Chanthaburi)
Euploea godarti [sic] ; Godfrey,1927 : 121. (N)
Euploea godarti [sic] ; Godfrey,1930 : 238. (all over)
Euploea layardi ; Godfrey,1930 : 238-239. (all over)
Euploea orontobates ; Godfrey,1930 : 239 (Chantharui)
Euploea core godartii ; Pinratana,1977 : 81, figs.D24b♂,♀. (except P)
Euploea core layardi ; Pinratana,1977 : 82, figs.D24c♂,♀. (P)
Euploea core godartii ; Pinratana & Eliot,1992 : 66,73, pl.81, fig.24a♂. (Bangkok; Chonburi; Chanthaburi; Kanchanaburi; Petchaburi; Ratchaburi)
Euploea core layardi ; Pinratana & Eliot,1992 : 66,73, pl.81, fig.24b♂. (Chanthaburi)
Euploea core godartii ; Yamaguchi & Aoki,2000 : 252-253, figs.61,♂,62,♂. (Mukdahan: Phu Sa Duak Bua; Phu Mu; Phu Huay Sing)
Euploea core godartii ; Yamaguchi & Aoki,2002 : 20. (Mukdahan: Ban Lao Kram; Phu Huay Sing; Nam Tok Tad Ton; Nam Tok Kham Cha I; Ban Na Lak; Ban Gok Tum; Ubon Ratchathani: Phu Chong Na Yoi; Nam Tok Soi Sawan; Kaeng E-Keow; Phu Hin Dang)
Euploea core godartii ; Yamaguchi, Aoki & Akiyama,2005 : 47. (Mukdahan: Ban Lao Kram; Phu Hin Khan; Ubon Ratchathani: Warin Chamrap; Nam Tok Soi Sawan)
Euploea core godartii ; Katsuyama et al.,2005 : 67, fig.104,♂. (Umphang: Tha-Sai; Umphang)
Euploea core godartii ; Katsuyama et al.,2006 : 48, fig.77,♂. (Nan: Ban Nam Poon)
Euploae core godartii ; Ek-Amnuay,[2007] : 202, pl.73, figs.D24a♂,♂(Un),♀,♀(Un),♀(f. layardi),♀(Un)(f. layardi) (Chiang Mai; Khao Yai; Pukae; Chanthaburi)
Euploae core godartii ; Ek-Amnuay,2012 : 202, pl.73, figs.N24a♂,♂(Un),♀,♀(Un),♀(f. layardi),♀(Un,f. layardi). (Chiang Mai; Khao Yai; Chanthaburi) (in part)
Euploae orontobates ; Ek-Amnuay,2012 : 204, pl.74, figs.N25,♂(Trat),♀(Sakerat),♀(Sakaerat,Un). (Nakhon Ratchasima; Chon Buri; Chanthaburi; Trat) (in part)
Euploae core godartii ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Euploea core godartii ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Chaiyaphum; Sa Kaeo; Chanthaburi) (accessed: 2023.07.16)
Euploae core godartii ; Kimura et al.,2016 : 36, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un)<♀,♀(Un). (Doi Chiang Dao; Phu Khiewo; Nakohn Nayork; Wan Nam Khiewo; Ban Na Lak; Kao Soi Dao; Kao Saverb; Kratching WF; Bang Phra; Kao Khiewo; Bangkok; Doi Musso; Sai York Noi; Kanchanaburi; Kaeng Krachang)
Euploae core godartii ; Chaianunporn & Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Euploea core godartii ; Motono & Negishi,1989 : 52, pl.35, figs.1,♂,2,♂(Un),3,♀,4,♀(Un). (Tha Ngone; Vientiane)
Euploea core gorartii ; Osada, Uémura & Uehara,1999 : 204, pl.44, figs.♂,♀. (Lak Sao; Luang Phrabang)
Euploae core layardi ; Osada, Uémura & Uehara,1999 : 204, pl.44, fig.♂. (Vientiane; Oudomxay)
Euploea core godartii ; Nakamura & Wakahara,2012 : 43.
Euploea core godartii f. layardi ; Nakamura & Wakahara,2012 : 43.
Euploea core godartii ; Onodera,2022 : 23, pl.83, figs.♂,♂(Un),♀,♀(Un),♂(f. layardi),♂(Un,f. layardi),♀(f. layardi),♀(Un,f. layardi). (Bolikhamxai: Thabok; Lak Sao; Khammouan: Natung; Nahin)

[ CAMBODIA ]
Euploea core godartii ; Woodfield & Murton,2006 : figs.♂,♂(Un). (Botum Sakor NP.)
[ Euploea sp. ; Woodfield & Murton,2006 : figs.♂,♂(Un). (Botum Sakor NP.) ]
Euploea core ; Onodera,2007 : 23, fig.26,♀. (Rattanakiri: Veun Sai)
Euploea core godartii ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)
Euploea core godarti [sic] ; Kosterin,2019 : 94. (Preah Sihanouk; Kep)
Euploea core godarti [sic] ; Kosterin,2019b : 30. (Ratanakiri)
Euploea core godarti [sic] ; Kosterin,2020 : 323. (Siem Reap; Stung Treng)
Euploea core godarti [sic] ; Chartier & Kosterin,2022 : 115. (Kep; Koh Kong; Kampot; Mondulkiri; Phnom Penh; Preah Sihanouk; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Euploea distanti circuita ; Fruhstorfer,1910 : 236. (Tonkin)
Euploea godarti godarti [sic] ; Fruhstorfer,1910 : 236. (Tonkin / Siam)
Euploea godarti [sic] ; Godfrey,1919 : 465. (S: Daban)
Euploea circuita ; Vitalis,1919 : 221. (Cochinchine)
Euploea tonkinensis ; Vitalis,1919 : 221. (Tonkin)
Euploea core f. core ; Dubois & Vitalis,1921 : 24. (Saigon; Cochinchine)
Euploea core ab.tonkinensis ; Dubois & Vitalis,1921 : 25. (Tonkin)
Euploea layardi ; Moulton,1923 : 128. (Pulo Condore)
Euploea godarti [sic] ; Lemée,1950 : 64. (Saigon)
Euploea core core ; Metaye,1957 : 101. (S)
Euploea distanti circuita ; Metaye,1957 : 101. (N / S)
Euploea godarti godarti [sic] ; Metaye,1957 : 101. (N / C / S)
Euploea godarti godarti [sic] f. dominans ; Metaye,1957 : 101. (N / C / S)
Euploea godarti godarti [sic] f. layardi ; Metaye,1957 : 101. (N / C / S)
Euploea godarti godarti [sic] ab. tonkinensis ; Metaye,1957 : 101. (N)
Euploea distanti f.circuita ; Metaye,1958a : 162-163. (N / S: Trang Bom)
Euploea alcathoe godarti [sic] ; Metaye,1958a : 163, pl.2, fig.6,♂. (all over)
Euploea alcathoe godarti [sic] f. dominans ; Metaye,1958a : 163. (all over)
Euploea alcathoe godarti [sic] f. layardi ; Metaye,1958a : 163. (all over)
Euploea alcathoe godarti [sic] ab. tonkinensis ; Metaye,1958a : 163. (N)
Euploea core godartii ; Morishita,1970 : 68, figs.19A♂,♀ ; 19B♂,♂ Variation forms. (Saigon; Cholon; Dinh Quan)
Euploea core godartii ; Ikeda et al.,1999 : 53, figs.2:9,♂,11,♂,12,♂(Un),13,♀,14,♀(Un),9:9(♂genitalia). (Cuc Phuong) (in part)
Euploea core ; Vu,2003 : 56. (Phu Quoc)
Euploea core godartii ; Luong et al.,2004 : 3, 39, figs.♂,♂(Un), 40, figs.♀,♀(Un). (Cuc Phuong)
Euploea core godartii ; Miyazaki, Saito & Saito,2006 : 15, figs.D-17,♂♀. (Lam Dong: Dambri; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Euploea core godartii ; Nihira,2009 : 98. (Phu Quoc)
Euploea core godartii ; Monastyrskii,2011 : 53-55, pl.7, figs.c♂(f. graminifera),d♀(f. graminifera),e♂(f. godartii),f♀(f. godartii). (N: Ba Be; Na Hang; Huu Lien; Ba Vi; Tam Dao; Cat Ba; Cuc Phuong / C: Xuu Lien; Ben En; Pu M123at[Pu Mat]; Vu Quang; Huong Son; Phong Nha-Ke Bang; Phong Dien; Dak Rong; Huong Thuy; Nam Dong; Bach Ma; Kon Cha Rang; Kon Ka Kinh; Hon Ba; Bao Lam / S: La Nga; Nghia Trung; Binh Chau; Cat Tien; Lo Go Sa Mat; Phu Quoc Is.)
Euploea core godartii ; Monastyrskii & Devyatkin,2015 : 19. (C / S)
Euploea core godartii ; Saito et al.,2017 : 80. (S: Nui Cam; Nui Coto; Nui Ket)

[ THAILAND ]
Mae Hong Son : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Rai ; 2♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 11♂ 5♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phrae ; 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 2♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Kanchanaburi : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchabun : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chaiyaphum : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nakhon Nayok : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chonburi : 1♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 3♂ 3♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Trat : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchaburi : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Vientiane : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ VIETNAM ]
Lao Cai : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Lam Dong : 4♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Kien Giang (Phu Quoc) : 4♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is common in secondary and montane forests at low to moderate elevations (Alt. 10 - 1000m). Both sexes visit flowers, and males occasionally come to the banks of streams and puddles to suck water.
This taxon is polymorphic and has very variable markings. There is a transition zone between graminifera in peninsular Thailand and amymone in northern Vietnam, where both subspecies occur together and intermediate forms are frequently observed. The name tonkinensis seems to be an intermediate form between godartii and amymone. The taxon f. layardi (binghami is the same population) and intermediate forms of layardi and godartii or graminifera are found throughout the region from north to south, sympatric with godartii and graminifera. However, the typical layardi is commonly observed in eastern to S.E.Thailand to S.Vietnam, while intermediate forms are more common in other areas.
All specimens shown in Ek-Amnuay, 2012 as Euploea orontobates are misidentifications of Euploea core godartii f. layardi. This species is indistinguishable from several similar species by the UnH markings. The androconial brand on the UnF and the UpF markings must be examined.
As I mentioned in the section on Euploea crameri praedicabilis, the type specimen of circuita Swinhoe, 1903, which is now considered a synonym of Euploea core, bears a striking resemblance to the branded crameri found in S.E. Thailand. Therefore, it is necessary to confirm the brand on the UnF of the type specimen.

Lucas, P.H.,1853 : Description de nouvelles especes de Lepidopteres appartenant aux collections entomologiques du Musee de Paris.
Revue Mag. Zool. (2)5:272-275, 310-322.

Felder, C. & Felder, R.,[1865]-1874 : Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair.
Zoologischer Theil. Zweiter Band. Zweite Abtheilung: Lepidoptera. Wien, Carl Gerold's Sohn.
(1):[4] + 1-136, pls. 1-21 ([12 January 1865]) ; (2):[2] + 137-378, pls. 22-47 ([after October] 1865) ; (3):[2] + 379-536, pls.48-74 ([25 April] 1867) ; (4):[6] + 1-9 [explanation of plates] ([31 December] 1874)

Druce, H.,1874 : A List of the Lepidopterous Insects collected by Mr. L. Layard at Chentaboon and Nahconchaisee, Siam,
with Descriptions of new Species.
Proc. zool. Soc. Lond. 1874(1):102-109, pl.16, 6 figs.

Moore, F.,1879 : A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim,
with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4):821-858, pls. 51-53.

Moore, F.,1883 :
A Monograph of Limnaina and Euploeina, two Groups of Diurnal Lepidoptera belonging to the Subfamily Euploeinae;
with descriptions of new genera and species. Part II. Euploeina.
Proc. zool. Soc. Lond. 1883(3):253-324, pls.29-32.

Swinhoe, C.,1903 : New species of Eastern and African Lepidoptera.
Ann. Mag. nat. Hist. (7)11:499-511.

Fruhstorfer, H.,1910 : 3. Familie: Danaidae in Seitz, Gross-Schmett. Erde 9
225-240. (15.Aug.1910)

Talbot, G.,1940 : Some new Forms of the Genus Euploea Fabr. (Lep., Danaidae)
Ent. mon. Mag. 76:247-248.

Murayama, S.,1961 : An unrecorded and some aberrant butterflies from Formosa
Trans. lepid. Soc. Jap. 11(4):54-61.

♂, Chiang Rai, Thailand.	♂, Chiang Mai, Thailand.

1998.11.15 - 2024.02.29.