Cyrestis cocles cocles

Cyrestis cocles cocles (Fabricius,1787)


♂, Xaignabouli, Laos.		♀, Champasak, Laos.

♂, Chanthaburi, Thailand.		♀, Phetchaburi, Thailand.

♂, Vinh Phuc, Vietnam.		♂, Chanthaburi, Thailand.

♂, Chanthaburi, Thailand.		♂, Chanthaburi, Thailand.

♂, Phetchaburi, Thailand.		♂, Prachuap Khiri Khan, Thailand.

ON : Papilio cocles Fabricius,1787
OD : Mantisa. Ins. 2:7.
TL : "Siam" (♂, NHML)
Distribution : India to Myanmar, Andamans, Thailand,
Laos, Cambodia, Vietnam, Yunnan, Hainan.

● ssp.cocles
● transition zone of ssp.cocles and earli
● ssp.earli

Synonyms

Cyrestis formosa Felder & Felder,[1867] : Reise Fregatte Novara, Lep. Rhop. (3):412. TL. Ins. Andaman.
Cyrestis cocles var.andamanica Wood-Mason & de Nicéville,1881 : J. asiat. Soc. Bengal, Pt2. 50(4):246. TL. Amdamans.
Cyrestis horatius Wood-Mason & de Nicéville,1881 : Proc. asiat. Soc. Bengal 1881:142. TL. S.Andamans.
Cyrestis natta Swinhoe,1899 : Ann. Mag. nat. Hist. (7)3:105. TL. Kasia Hills.
Cyrestis cocles cocleoides Fruhstorfer,[1902] : Deut. ent. Zeit. [Iris] 14(2):274. TL. Than-Moi, Tonkin ; Xom-Gom, S.Annam.

Type Material Information

andamanica ; Type ♂, Andaman. (NZC) ; Sheela et al.,2019 : 188, fig.♂,♂(Un).

Records taken from Literature

[ THAILAND ]
Cyrestis cocles cocles ; Godfrey,1916 : 127-128. (Petchaburi River; Chonburi: Hup Bon)
Cyrestis cocles cocles ; Godfrey,1927 : 123. (N)
Cyrestis cocles cocles ; Godfrey,1930 : 276. ("Prae: Me Song forest"; "Chiang Rai"; Muak Lek; Petchaburi River) (in part)
Cyrestis cocles cocles ; Pinratana,1979 : 37, figs.N53a♂(WSF),♂(Un,WSF),♂(DSF),♂(Un,DSF). (Chanthaburi; Loei; Petchaburi; Uthai Thani; Kanchanaburi; Nakhon Nayok; Prachuap Khiri Khan)
Cyrestis cocles cocles ; Pinratana & Eliot,1996 : 42, pl.24,figs.55,♂,♂(f.earli),♂(f.cocles). (Nakhon Ratchasima; Nakhon Nayok; Chonburi; Chanthaburi; Kanchanaburi; Petchaburi) (in part)
Cyrestis cocles cocles ; Yamaguchi & Aoki,2000 : 254-255, fig.93,♂. (Mukdahan: Phu Pha Thaep)
Cyrestis cocles cocles ; Yamaguchi & Aoki,2002 : 27. (Ubon Ratchathani: Ban Na Pho Krang; Mukdahan: Ban Lao Kram; Ban Na Lak; Ban Gok Tum)
Cyrestis cocles cocles ; Yamaguchi, Aoki & Akiyama,2005 : 50. (Ubon Ratchathani: Nam Tok Soi Sawan; Mukdahan: Ban Lao Kram; Phu Huai Sing)
Cyrestis cocles cocles ; Ek-Amnuay,[2007] : 384, pl.166, fig.N57a♂. (Nakhon Nayok)
Cyrestis cocles earli ; Ek-Amnuay,[2007] : 384, pl.166, figs.N57b♂(center),♂(right). (in part)
Cyrestis cocles cocles ; Ek-Amnuay,2012 : 478, pl.212,fig.N340a♂. (Nakhon Nayok; Khao Soi Dao)
Cyrestis cocles earli ; Ek-Amnuay,2012 : 478, pl.212,figs.N340b♂(center),♂(right). (in part)
Cyrestis cocles cocles ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Cyrestis cocles cocles ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Khon Kaen; Nakhon Ratchasima; Sa Kaeo; Chonburi; Phetchaburi) (accessed: 2021.07.20)
Cyrestis cocles cocles ; Kimura et al.,2016 : 117, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un); 118, figs.♀,♀(Un),♀,♀(Un),♀,♀(Un). (Sarika; Wan Nam Khiewo; Kao Soi Dao; Kae Khiewo; Sai York Noi; Erawan WF; Kao Phang WF; Kaeng Krachan)

[ LAOS ]
Cyrestis cocles f.cocles ; Dubois & Vitalis de Salvaza,1924 : 38. (Laos)
Cyrestis cocles f.cocleoides ; Dubois & Vitalis de Salvaza,1924 : 38. (Laos)
Cyrestis cocles cocles ; Motono & Negishi,1989 : 56, pl.53, figs.1,♂,2,♂(Un),3,♂,4,♂(Un). (Vang Vieng; Phatong; Tha Ngone; Vientiane; Pakse; Luang Prabang; Nam Ngum Dam)
Cyrestis cocles cocles ; Osada, Uémura & Uehara,1999 : 206, pl.51, figs.♂(f.earli),♀(f.cocles). (Vang Vieng; Thakhek)
Cyrestis cocles cocles ; Nimura & Wakahara,2005 : 42. (Thakhek)
Cyrestis cocles cocles ; Onodera,2009b : 35,39. (Vientiane: Namsong Riverside; Bolikhamxai: Thabok)
Cyrestis cocles cocles ; Nakamura & Wakahara,2012 : 45.

[ CAMBODIA ]
Cyrestis cocles ; Onodera,2007 : 25, figs.49♂,50,♀. (Rattanakiri: Ou Chum)
Cyrestis cocles ; Onodera,2009 : 67,68. (Siem Reap: Phnum Kulen)
Cyrestis cocles cocles ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)

[ VIETNAM ]
Cyrestis cocles ; Joannis,1901 : 320. (Tonkin: Cao Bang)
Cyrestis cocles cocleoides ; Fruhstorfer,1912 : 574. (C.Tonkin / S.Annam)
Cyrestis cocleoides ; Dubois & Vitalis de Salvaza,1919 : 228. (Tonkin)
Cyrestis cocles f.cocles ; Dubois & Vitalis de Salvaza,1924 : 38. (Tonkin)
Cyrestis cocles f.cocleoides ; Dubois & Vitalis de Salvaza,1924 : 38. (Tonkin / Annam)
Cyrestis cocles f.earli ; Dubois & Vitalis de Salvaza,1924 : 38. (Cochinchine)
Cyrestis cocles ; Lemee,1950 : 10. (Hagiang)
Cyrestis cocles cocles ; Metaye,1957 : 95. (N / C)
Cyrestis cocles cocleoides ; Metaye,1957 : 95. (N / C)
Cyrestis cocles earli ; Metaye,1957 : 95. (S)
Cyrestis cocles cocles ; Ikeda et al.,2000 : 30, figs.7:17,♂,18,♂(Un),19,♀,20,♀, fig.9:20(♂,genitalia). (Cuc Phuong)
Cyrestis cocles cocles ; Hao et al.,2004 : 6, 61, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Cyrestis cocles ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Phong Dien)
Cyrestis cocles cocles ; Miyazaki, Saito & Saito,2006b : 20, figs.N-36,♂,♀. (Lam Dong: Dai Duc Me; Bin Thuan: Dong Tien)
Cyrestis cocles cocles ; Monastyrskii & Devyatkin,2015 : 34. (N / C / S)
Cyrestis clcles cocles ; Monastyrskii,2019 : 4(1):144-145, pl.32, fig.1a♂,1b♂,1c♂(Un). (N: Tam Dao; Ba Be; Na Hang; Huu Lien; Cat Ba; Cuc Phuong / C: Pu Mat; Vu Quang; Phong Nha Ke Bang; Phong Dien; Son Thanh; Nui Chua; Chu Yang Sin / S: Nam Cat Tien; La Nga; Lo Go Xa Mat; Binh Chau)

Specimens Examined

[ THAILAND ]
Kanchanaburi : 3♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Sa Kaeo : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chanthaburi : 6♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phetchaburi : 3♂ 1♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Prachuap Khiri Khan : 4♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ LAOS ]
Xaignabouli : 2♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Khammouan : 1♀ (photo by H.Onodera).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Champasak : 1♀.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ VIETNAM ]
Vinh Phuc : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Notes
The butterfly is locally not rare in montane forest at low elevations (alt.200m - 450m) and often rest on the underside of leaves with their wings spread.
Godfrey,1930 recorded it from Chiang Rai and Phrae, N.Thailand, however are dubious. Further distribution research in northern Thailand is needed. The specimen from Phetchaburi to Prachuap Khiri Khan seems intermediate between ssp.cocles and ssp.earli, it seems transition zone of both subspecies.
Corbet,1941 assumed the type locality of cocles as Pulau Salang [Phuket], Siam. If so, the population in Phuket is most likely to be the same subspecies as Malay Peninsula subspecies earli, then ssp.earli should be the synonym of cocles, and the continental populations should be given an alias. Further taxonomic study is needed.

Literature

Fabricius, J.C.,1787 : Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis,
differentiis specificis, emendationibus, observationibus.
Copenhagen, Christ. Gottl. Proft 8vo 1:xx+348pp ; 2:[2]+382pp

Felder, C. & R. Felder,1865-1875 :
Rhopalocera. In Reise der Osterreichischen Fregatte "Novara" um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Zweiter Band:
Abtheilung. Wien : vi + 548 + [1] pp in 5 parts, 140 pls.

Wood-Mason, J. & C.L.A. de Nicéville,1881 : Abstract of: Second List of Rhopalocerous Lepidoptera from the Andaman Islands, with Descriptions of new or little-known species and Varieties.
Proc. asiat. Soc. Bengal 1881:142-143.

Wood-Mason, J. & C.L.A. de Nicéville,1881 : Second List of Rhopalocerous Lepidoptera from Port Blair, Andaman Island.
J. asiat. Soc. Bengal, Pt.2. 50(4):243-262, pl.14.

Swinhoe, C.,1899 : New Species of Orientali Lepidoptera.
Ann. Mag. nat. Hist. (7)3:102-116.

Fruhstorfer, H.,[1902] : Neue und seltene Lepidopteren aus Annam und Tonkin und dem malayischen Archipel.
Deut. ent. Zeit. [Iris] 14(2):265-276, pl.5.

Corbet, A.S.,1941 : Observations on Certain of the Fabrician Names onf Indo-Australian Rhopalocera (Lepid.)
Proc. R. ent. Soc. Lond. (B)10(6):98-106.


♂, Sa Kaeo, Thailand.	♂, Kanchanaburi, Thailand.

2007.10.10 - 2021.08.31.