Euripus nyctelius nyctelius  (Doubleday,1845)

Euripus nyctelius nyctelius ♂ Up. Euripus nyctelius nyctelius ♂ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♂, Mae Hong Son, Thailand. ♀, Chiang Mai, Thailand.
Euripus nyctelius nyctelius ♂ Up. Euripus nyctelius nyctelius ♂ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Euripus nyctelius nyctelius ♂ Up. Euripus nyctelius nyctelius ♂ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♂, Nan, Thailand. ♀, Nan, Thailand.
Euripus nyctelius nyctelius ♂ Up. Euripus nyctelius nyctelius ♂ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♂, Prachuap Khiri Khan, Thailand. ♀, Nan, Thailand.
Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♀, Phrae, Thailand. ♀, Tak, Thailand.
Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un. Euripus nyctelius nyctelius ♀ Up. Euripus nyctelius nyctelius ♀ Un.
♀, Prachuap Khiri Khan, Thailand. ♀, Lampang, Thailand.

ON : Diadema nyctelius Doubleday,1845
OD : Ann. Mag. nat. Hist. (1)16:182.
TL : Sylhet.

Distribution :
N.India, Assam to Myanmar, Thailand, Laos, Vietnam, S.China, Hainan.

Synonyms

Euripus halitherses Doubleday,[1848] : Gen. diurn. Lep. (2):293, pl.41, fig.2. TL. N.India : Assam.
Hestina isa Moore,[1858] : in Horsfield & Moore, Cat. Lep. Mus. East India Co. (1):161. TL. Darjeeling. (♀)
Euripus haliartus Felder & Felder,1864 : Wien. ent. Monats. 4(8):234-235. TL. Sikkim. (♀)
Euripus cinnamomeus Wood-Mason,1881 : J. asiat. Soc. Bengal, Pt.2, 50(4):272-273, pl.4, fig.4♀. TL. Shilong, Khasi Hills. (♀)
Euripus halitherses ♀f.alcathoeides de Nicéville,1886 : Butts. India Burmah Ceylon 2:20. TL. Tenasserim.
Euripus halitherses mastor Fruhstorfer,1903 : Deut. ent. Zeit. [Iris] 16:28-33. TL. Tonkin.
Euripus halitherses ♀ab.neda Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:29. TL. Chiem-Hoa, Tonkin.
Euripus halitherses ♀ab.phygalia Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:29. TL. Chiem-Hoa, Tonkin.
Euripus halitherses ♀ab.hadria Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:30. TL. Chiem-Hoa, Tonkin.
Euripus halitherses ♀ab.pademoides Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:30. TL. Sikkim.
Euripus halitherses ♀ab.gulussa Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:30. TL. Sikkim.
Euripus halitherses ♀ab.gyrtone Fruhstofer,1903 : Deut. ent. Zeit. [Iris] 16:30-31. TL. Sikkim.
* ♀v.isis Rosa,1929 : TL. Karen Hill.
* ♀f.pseuduniformis Nguyen,1980 : TL. Lower Burma.
Euripus nyctelius marginalis Chou, Yuan, Yin, Zhang & Chen,2002 : Entomotaxonomia 24(1):53. TL. Guangdong.


Records taken from Literature

Euripus halitherses mastor ♀ab.isa ; Fruhstorfer,1906 : 318. (Siam)
Euripus halitherses mastor ♀ab.neda ; Fruhstorfer,1906 : 318. (Siam)
Euripus halitherses mastor ♀ab.alcathoeoides ; Fruhstorfer,1906 : 318. (Hinlap)
Euripus halitherses mastor ; Fruhstorfer,1914 : 708. (Siam)
Euripus halitherses halitherses ; Godfrey,1916 : 132.
Euripus halitherses halitherses ; Godfrey,1930 : 299-300. (Chiang Rai / Nam Phat(Sen Tor) / Muak Lek)
Euripus halitherses mastor ; Davidson & Macbeth,1938 : 75. (Lampang / Chiang Mai: Doi Suthep)
Euripus nyctelius euploeoides ; Pinratana,1979 : 94, figs.N161,♂,♀,♀,♀,♀,♀,♀. (Chiang Mai / Chaiyaphum / Chanthaburi / Nakhon Nayok / Prachinburi) (in part)
Euripus nyctelius nyctelius ♀f.nyctelius ; Pinratana & Eliot,1996 : 90, pl.67. (Nakhon Ratchasima / Chiang Mai / Chaiyaphum / Chiang Rai / Phrae / Phisanulok / Nakhon Nayok / Chanthaburi / Petchaburi) (in part)
Euripus nyctelius nyctelius ♀f.euploeoides ; Pinratana & Eliot,1996 : 90, pl.67. (Chaiyaphum / Nakhon Ratchasima / Phrae)
Euripus nyctelius nyctelius ♀f.alcathoides ; Pinratana & Eliot,1996 : 90, pl.67. (Phrae)
Euripus nyctelius euploeoides ; Yamaguchi & Aoki,2000 : 256, fig.114,♂. (Ubon Ratchathani: Pha Taem / Mukdahan: Phu Pha Thaep ; Phu Pha Phung)
Euripus nyctelius euploeoides ; Yamaguchi & Aoki,2002 : 31. (Ubon Ratchathani: Kaeng E-Keow)
Euripus nyctelius euploeoides ; Yamaguchi, Aoki & Akiyama,2005 : 52. (Mukdahan: Ban Lao Kram)
Euripus nyctelius euploeoides ; Katsuyama et al.,2005 : 70, fig.174,♂. (Umphang)
Euripus nyctelius euploeoides ; Katsuyama et al.,2006 : 50, fig.123,♂. (Nan: Ban Nam Poon)
Euripus nyctelius nyctelius ; Ek-Amnuay,[2007] : 470, pl.209, figs.N185, ♂,♂(Un),♀f.cinnamomea,♀f.alcathoeoides, ♀f.euploeoides,♀f.nyctelius,♀f.isa. (Chiang Mai / Nakhon Nayok)
Euripus nyctelius nyctelius ; Ek-Amnuay,2012 : 492, pl.219, figs.N369, ♂,♂(Un),♀f.cinnamomea,♀f.alcathoeoides, ♀f.euploeoides,♀f.nyctelius,♀f.isa. (Chiang Mai / Nakhon Nayok)
Euripus nyctelius nyctelius ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs, Early Stages. (Chaiyaphum / Kamphaeng Phet / Phetchabun / Chiang Mai / Nakhon Nayok / Sa Kaeo)
Euripus nyctelius nyctelius ; Kimura et al.,2016 : 126-127, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Doi Pha Hom Pok / Doi Suthep / Khun Lao Villege / Bang Salock / Mae Mo / Doi Pa Chang / Thung Saleng Luang / Phu Khiewo / Wan Nam Khiewo / Kratchin WF / Kao Khiewo / Kaeng Krachan) (in part)
Euripus nyctelius nyctelius ; Chaianunporn & Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Euripus cinnamomeus ; Dubois & Vitalis de Salvaza,1919 : 233. (Laos)
Euripus nyctelius ssp. ; Motono & Negishi,1989 : 62, pl.76, figs.5,♂,6,♂(Un),7,♀,8,♀. (Luang Prabang / Pakse / Tha Ngone / Nam Moh / Vientiane / Ban Houei Sai / Nam Ngum Dam)
Euripus nyctelius euploeoides ; Nimura & Wakahara,2005 : 43. (Thabeng / Namdhua / Nakai)
Euripus nyctelius euploeoides ; Nakamura et al.,2009 : 26, figs.10,♀,♀(Un). (Bolovens)
Euripus nyctelius euploeoides ; Onodera,2009b : 41. (Khammouan: Nacay)
Euripus nyctelius euploeoides ; Nakamura & Wakahara,2012 : 47, pl.6, figs.168,♀,♀(Un).

[ VIETNAM ]
Euripus halitherses mastor ; Fruhstorfer,1906 : 317. (Tonkin: Chiem Hoa)
Euripus halitherses mastor ♀ab.cinnamomeus ; Fruhstorfer,1906 : 318. (Tonkin: Phunkte / Annam)
Euripus halitherses mastor ♀ab.hadria ; Fruhstorfer,1906 : 318. (Tonkin)
Euripus halitherses mastor ♀ab.neda ; Fruhstorfer,1906 : 318. (Tonkin)
Euripus halitherses mastor ; Fruhstorfer,1914 : 708. (Tonkin)
Euripus halitherses mastor ♀ab.phygalia ; Fruhstorfer,1914 : 708. (Annam)
Euripus halitherses mastor ♀ab.cinnamomeus ; Fruhstorfer,1914 : 708. (Annam)
Euripus mastor ; Dubois & Vitalis de Salvaza,1919 : 233. (Tonkin)
Euripus phygalia ; Dubois & Vitalis de Salvaza,1919 : 233. (Tonkin)
Euripus neda ; Dubois & Vitalis de Salvaza,1919 : 233. (Tonkin)
Euripus badria ; Dubois & Vitalis de Salvaza,1919 : 233. (Annam)
Euripus mastor ; Lemee,1950 : 16. (Hagiang)
Euripus halitherses halitherses ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.neda ; Metaye,1957 : 98. (N)
Euripus halitherses halitherses f.♀.pademoides ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.haliartus ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.alcathoeoides ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.cinnamomeus ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.nyctelius ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses halitherses f.♀.gyrtone ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses mastor ; Metaye,1957 : 98. (N / C / S)
Euripus halitherses mastor f.♀.hadria ; Metaye,1957 : 98. (C)
Euripus nyctelius ; Hao et al.,2004 : 8, 70, figs.♂,♂(Un). (Cuc Phuong)
Euripus nyctelius ; Monastyrskii et al.,2006 : 42. (Thua Thien Hue: Nam Dong; A Luoi)
Euripus nyctelius mastor ; Miyazaki, Saito & Saito,2006b : 34, fig.N-109&,#9794;. (Lam Dong: Dambri)
Euripus nyctelius mastor ; Miyazaki, Saito & Saito,2007c : 38, fig.N-109&,#9792;. (Lam Dong: Dai Duc Me)
Euripus nyctelius mastor ; Monastyrskii & Devyatkin,2015 : 44. (N / C / S)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 12♂ 12♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 1♂ 1♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♀ (no data).
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 7♂ 10♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Uttaradit : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nongkhai : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 2♂ 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 7♂ 3♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 4♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Prachuap Khiri Khan : 12♂ 8♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Oudomxai : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Luang Prabang : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Phon Si : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Champasak : 1♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 2♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
Literature

Doubleday, E.,1845 : Descriptions of new or imperfectly described Diurnal Lepidoptera.
Ann. Mag. nat. Hist. (1)16:176-182, 232-236, 304-307.

Doubleday, E. & W.C. Hewitson,1846-1852 : The genera of diurnal Lepidoptera,
comprising their generic characters, a notice of their habits and transformations, and a catalogue of the species of each genus ;
illustrated with 86 plates by W.C.Hewitson.
Longon, Longman, Brown, Green and Longman's. xii + ii + 534 pp, 86 pls, In 54 parts. In 2 vols.

Horsfield, T. & F. Moore,[1858] :
A catalogue of the Lepidopterous Insects in the Museum of the Honourable East-India Company. Vol.1.
London. East India Co.

Felder, C. & R. Felder,1860 : Lepidopterologische Fragmente. VI.
Wien. ent. Monats. 4(8):225-251, pl.3,4.

Wood-Mason, J.,1881 : Description of a new Species of the Lepidopterous Genus Euripus from North-Eastern India.
J. asiat. Soc. Bengal, Pt.2. 50(4):272-273.

de Nicéville, C.L.A.,1886 : The butterflies of India, Burmah and Ceylon.
A descriptive handbook of all the known species of rhopalocerous Lepidoptera inhabiting that region, with notices of allied species occurring in the neighbouring countries along the border, with numerous illustrations. Volume II (Nymphalidae, Libytheidae, Nemeobiidae).
Calcutta Central Press : vii + 332 pp, frontispiece, pls.18-24.

Fruhstorfer, H.,1903 : Neue Nymphaliden aus Ostasien.
Deut. ent. Zeit. [Iris] 16(1):22-42.

Masui, A. & T. Inomata,1995 : Apaturinae of the World (Lepidoptera, Nymphalidae) 7. (in Japanese)
Yadoriga 160:2-16.

Lang, S-Y.,2012 : The Nymphalidae of China (Lepidoptera, Rhopalocera), Part I. Libytheinae, Danainae, Calinaginae, Morphinae, Heliconiinae, Nymphalinae, Charaxinae, Apaturinae, Cyrestinae, Biblidinae, Limenitinae.
Tshikolovets. Czech Rep. 456pp. col.pls.32.

Euripus nyctelius nyctelius ♂ Euripus nyctelius nyctelius ♀
♂, Phetchabun, Thailand. ♀, Chiang Mai, Thailand.
Euripus nyctelius nyctelius ♂ Euripus nyctelius nyctelius ♀
♂, Phetchabun, Thailand. ♀, Chiang Mai, Thailand.
Euripus nyctelius nyctelius ♂ Euripus nyctelius nyctelius ♀
♂, Chiang Mai, Thailand. ♀, Phetchabun, Thailand.
Euripus nyctelius nyctelius ♂ Euripus nyctelius nyctelius ♀
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.

2007.09.20 - 2020.03.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © Y.Inayoshi.