♂ (dsf), Phrae, Thailand.		♀ (dsf), Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Phong Saly, Laos.		♂, Luang Pabang, Laos. (photo by H.Onodera)
♂, Yala, Thailand.		♀, Yala, Thailand.
♂, Phetchabun, Thailand.		♀ (dsf), Chiang Mai, Thailand.

ON : Yphthima [sic] savara Grose-Smith, 1887 OD : Ann. Mag. nat. Hist. (5)20:267. TL : Burmah, Siamese frontier. (NHML) Distribution : Myanmar, Thailand, Laos, Vietnam, W.Malaysia.

Ypthima tonkiniana Fruhstorfer,1908 : Ent. Zeit. 22(31):127. TL. Chiem Hoa, N.Vietnam. (NHML)

savara ; Syntype 1♂, Rh3410, 1♂, Rh3411, Burmah, Siamese frontier. (NHML)
tonkiniana ; Syntype 1♀, #141995, Aug.Sept. Chiem Hoa, Tonkin. (NHML)

[ THAILAND ]
Ypthima savara savara ; Godfrey,1927 : 121. (N)
Ypthima savara savara ; Godfrey,1930 : 244. (Chiang Rai: Doi Chang; Khun Than; Prae: Me Song forest; Umphang)
Ypthima savara savara ; Aoki & Uémura,1984 : 83. figs.11,a-b. (Mae Hong Son: Mae Hong Son; Chiang Mai: Tha Tong; Tam Tab Tao; Chiang Dao; Mae Sa; Muang Ngai; Nam Tok Mae Klang; Chiang Rai: Wiang Pa Pao; Tak: Doi Musso; Kanchanaburi: Sai Yok Noi; Phitsanulok: Tong Salang Luang; Loei: Phu Kadung; Nam Nao)
Ypthima savara savara ; Pinratana,1988 : 33, pl.34, figs.92,♂,♀(Un). (Chiang Mai; Loei; Phitsanulok; Nakhon Ratchasima; Kanchanaburi; Tak)
Ypthima savara savara ; Yamaguchi & Aoki,2000 : 254, fig.79,♂(Un). (Mukdahan: Phu Pha Thaep)
Ypthima savara savara ; Yamaguchi & Aoki,2002 : 24. (Mukdahan: Ban Lao Kram; Ban Na Lak; Phu Huay Sing; Ban Gok Tum; Ubon Ratchathani: Ban Na Pho Krang; Phu Chong Na Yoi)
Ypthima savara savara ; Yamaguchi, Aoki & Akiyama,2005 : 49. (Mukdahan: Ban Lao Kram; Ban Na Lak; Ubon Ratchathani: Nam Tok Soi Sawan)
Ypthima savara savara ; Katsuyama et al.,2005 : 68, fig.128,♂. (Umphang: Tha-Sai)
Ypthima savara savara ; Katsuyama et al.,2006 : 49, fig.98,♂. (Nan: Ban Nam Poon)
Ypthima fasciata torone ; Ek-Amnuay,[2007] : pl.121, figs.S94,♂(top-left),♂(top-centerUn),♀(bottom),♀(bottom,Un). (in part)
Ypthima savara savara ; Ek-Amnuay,[2007] : 298, pl.121, figs.S96,♂,♂(Un),♀[♂],♀[♂](Un). (Chiang Mai; Phu Khieo)
Ypthima savara savara ; Ek-Amnuay,2012 : 298, pl.121, figs.N131,♂,♂(Un),♂(Un),♀,♀(Un). (Chiang Mai; Phu Khieo; Kaeng Krachan)
Ypthima savara savara ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Ypthima savara savara ; Kimura et al.,2016 : 194, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao; Doi Suthep; Thung Saleng Luang; Phu Kradung; Phu Khiewo; Wan Nam Khiewo; Ban Na Lak; Kratching WF; Kaeng Chai; Kao Phang WF; Sai York Noi; Kuwai Yai; Khlong Naka)
Ypthima savara savara ; Saito & Inayoshi,2018 : 30, figs.21,♂,♂(Un),22,♀,♀(Un). (Yala)
Ypthima savara savara ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Ypthima savara savara ; Motono & Negishi,1989 : 66, pl.91, figs.7,♂,8,♂(Un). (Vang Vieng)
Ypthima sarava savara ; Osada, Uémura & Uehara,1999 : 211, pl.96, figs.♂,♀(Un). (Thabok; Vientiane)
Ypthima sarava tonkiniana ; Uémura,2005 : 1, figs.4(map), 33,♂(wsf),34,♂(dsf). (Boun Neua; Thabok; Vieng Thong; Ban Boung Kuang; Lak Sao)
Ypthima sarava tonkiniana ; Uémura,2006 : 7, fig.39,map. (Vieng Thong)
Ypthima sarava tonkiniana ; Uémura,2007 : 3, fig.21,map. (Nakhai to Lak Sao)
Ypthima savara savara ; Onodera,2009b : 41. (Khammouan: Nacay)
Ypthima sarava tonkiniana ; Uémura,2011 : 3. (C: Phon Si; Nahin Pass; Nakhai to Lak Sao 20km; near Nam Teun)
Ypthima sarava tonkiniana ; Nakamura & Wakahara,2012 : 49.
Ypthima sarava tonkiniana ; Onodera,2022 : 61, pl.181, figs.♂(wsf),♂(wsf,Un),♂(dsf),♂(dsf,Un),♀,♀(Un). (Bolikamxai: Thabok)

[ CAMBODIA ]
Ypthima savara f.savara ; Dubois & Vitalis,1924 : 27. (Pnom-Penh)
Ypthima savara savara ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)
Ypthima savara ; Chartier & Kosterin,2022 : 116. (Koh Kong; Phnom Penh; Pursat)

[ VIETNAM ]
Ipthima [sic] tonkiniana ; Vitalis,1919 : 222. (Tonkin)
Ypthima savara f.tonkiniana ; Dubois & Vitalis,1924 : 27. (Tonkin: Chiem-Hoa)
Ypthima savara tonkiniana ; Metaye,1957 : 102. (N)
Ypthima savara savara ; Uémura & Monastyrskii,2004 : 24-25, figs.2i, 5.1,♂. (C: Phong Dien; Bach Ma; Ba Na; Kon Ka Kinh; Ban Don; Bao Loc / S: Nghia Trung; Cat Tien)
Ypthima savara tonkiniana ; Uémura & Monastyrskii,2004 : 24, figs.2g-h, 5.2,♂. (N: Ba Be / C: Ben En; Pu Mat)
Ypthima savara savara ; Monastyrskii,2005 : 110-111. (C: Phong Dien; Bach Ma; Kon Ka Kinh; Ban Don; Cat Tien; Da Teh / S: La Nga; Nghia Trung)
Ypthima savara tonkiniana ; Monastyrskii,2005 : 110-111, pl.29, figs.1a♂,1b♂(Un),1c♀,1d♀(Un). (N: Na Hang; Ba Be / C: Ben En; Pu Mat)
Ypthima savara savara ; Miyazaki, Saito & Saito,2006c : 34, fig.S-33,♀[♂]. (Lam Dong: Dambri)
Ypthima savara savara ; Miyazaki, Saito & Saito,2007c : 39, fig.S-33,♀. (Lam Dong: Dambri)
Ypthima savara savara ; Monastyrskii & Devyatkin,2015 : 26. (C: to Lam Dong)
Ypthima savara tonkiniana ; Monastyrskii & Devyatkin,2015 : 26. (N / C: to Quang Tri)

[ THAILAND ]
Mae Hong Son : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 11♂ 11♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phrae : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Loei : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchabun : 1♂ 1♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chaiyaphum : 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchaburi : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Yala : 4♂ 1♀ (incl. photo by S.Atdhabhan).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Luang Phabang : 1♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phong Saly : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Oudomxai : 1♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Bolikhamxai : 2♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Vientiane : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Sekong : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is common in montane forests at low to moderate elevations (Alt. 200 - 1100m).

Grose-Smith, H.,1887 : Descriptions of eight new Species of Asiatic Butterflies.
Ann. Mag. nat. Hist. (5)20:265-268.

Fruhstorfer, H.,1908 : Neue ostasiatische Satyriden
Ent. Zeit. 22(31):127.

Uémura, Y.,2020 : A synonymic list of the sakra-group of the genus Ypthima Hübner (Lepidoptera: Nymphalidae, Satyrinae)
Bull. Toyosato Mus. Ent. (30):1-20.

♂ (dsf), Chiang Mai, Thailand.	♀ (dsf), Phetchabun, Thailand.
♂, Chiang Mai, Thailand.	♂+♀ (mating), Chiang Mai, Thailand.
♂ (dsf), Chiang Mai, Thailand.	♂ (dsf), Yala, Thailand. (photo by S.Atdhabhan)

1998.12.08 - 2023.07.31.