Graphium doson actor  Fruhstorfer, 1907

Graphium doson actor ♂ Up. Graphium doson actor ♂ Un. Graphium doson actor ♀ Up. Graphium doson actor ♀ Un.
♂, Chiang Mai, Thailand. (photo by A.M.Cotton) ♀, Chiang Mai, Thailand.
Graphium doson actor ♂ Up. Graphium doson actor ♂ Un. Graphium doson actor ♀ Up. Graphium doson actor ♀ Un.
♂, Chanthaburi, Thailand. ♀, Nan, Thailand.
Graphium doson actor ♂ Un. Graphium doson actor ♂ Un. Graphium doson actor ♀ Up. Graphium doson actor ♀ Up.
♂(un), Chanthaburi, Thailand. ♂ (un), Mae Hong Son, Thailand. ♀, Kanchanaburi, Thailand. ♀, Phrae, Thailand.
Graphium doson actor ♂ Up. Graphium doson actor ♂ Un. Graphium doson actor ♀ Up. Graphium doson actor ♀ Un.
♂, Dong Nai, Vietnam. (photo by K.Saito) ♀, Dong Nai, Vietnam. (photo by K.Saito)

ON : Graphium doson actor Fruhstorfer, 1907
OD : Ent. Zeit. 21(34):209.
TL : Hainan. (♂, NHML)
Distribution : Pakistan, Nepal, India to Bangladesh, Myanmar,
Thailand, Laos, Cambodia, Vietnam, Yunnan, Guangdong, Hainan,
Hong Kong.
● ssp.actor
ssp.robinson
ssp.evemonides
Graphium doson actor Map

Synonyms

Papilio axion Felder & Felder, 1864 : Verh. zool.-bot. Ges. Wien. 14:305. TL. 'India sept.' [Silhet]. (homonym)
Papilio jason axion gen.vern.nivepictus Fruhstorfer, 1903 : Berl. ent. Z. 47(3/4):212. TL. Sikkim ; Siam ; Assam ; Hainan.
Papilio eurypylus priscus ab.aurifer Fruhstorfer, 1907 : Ent. Zeit. 21(34):182. TL. Siam. (♂, NHML)
Papilio jason nanus Fruhstorfer, 1909 : Ent. Zeit. 22(41):170. TL. Xom-Gom, S.Annam. (♂, NHML)
Papilio jason f.praestabilis Fruhstorfer, 1909 : Ent. Zeit. 22(41):170. TL. Tonkin ; Hong Kong. (♂, NHML)
Graphium choui Huang, 2001 : in Jiang, Huang & Yao, Fauna of insects in Fujian Province of China 4:23,152, pl.26-47A♀, B♀(Un). TL. Fujian. (♀, Fujian Agricultural University)
Graphium doson axionides Page & Treadaway, 2014 ; Stutt. Beit. Natur. Serie A, Neue Ser. 7:272-273. TL. India sept. (Silhet). Replacement name for Papilio axion (Felder & Felder)


Type Materials Information

actor ; ♂, Type HT (red), #149390, Hainan, Whitehead (NHML, examined)
aurifer ; ♂, Type (red), #149391, Jan. Hinlap, Siam. H.Fruhstorfer (NHML, examined)
nanus ; ♂, Type HT (red), #149389, Februar, Xom-Gom, Sud-Annam. H.Fruhstorfer (NHML, examined)
praestabilis ;
♂, Type HT (red), #149387, Aug.Stpt., Chiem-Hoa, Central-Tonkin. H.Fruhstorfer (NHML, examined)
♀, Type (red), #149388, Aug.Stpt., Chiem-Hoa, Central-Tonkin. H.Fruhstorfer (NHML, examined)
choui ; Holotype ♀, 12.Jun.1992. Shunchang, Fujian, China. L.Yao (Fujian Agricultural University)


Records taken from Literature

[ THAILAND ]
Papilio jason axion ; Fruhstorfer,1903 : 204-206. (Hinlap; Kanburi)
Papilio doson axion ; Jordan,1909 : 97. (Siam)
Papilio doson axion ; Godfrey,1916 : 112.
Papilio doson axion ; Godfrey,1927 : 120. (N)
Papilio doson axion ; Godfrey,1930 : 217-218. (all over)
Papilio doson praestabilis ; Davidson & Macbeth,1938 : 69-70. (N)
Graphium doson axion ; Pinratana,1977 : 56, figs.P44a♂,♂(Un). (all over)
Graphium doson axion ; Pinratana & Eliot,1992 : 35,46, pl.65, figs.56a♀,♂(Un). (Nakhon Nayok; Kanchanaburi)
Graphium doson axion ; Yamaguchi & Aoki,2000 : 251, fig.42,♂. (Mukdahan: Phu Mu)
Graphium doson actor ; Kocak & Kemal,2000 : 4. (Thailand)
Graphium doson axion ; Yamaguchi & Aoki,2002 : 16. (Mukdahan: Ban Lao Kram; Phu Huay Sing; Ban Na Lak; Ban Gok Tum; Ubon Ratchathani: Nam Tok Soi Sawan; Kaeng E-Keow)
Graphium doson axion ; Yamaguchi, Aoki & Akiyama,2005 : 46. (Ubon Ratchathani: Phu Chong Na Yoi; Mukdahan: Ban Lao Kram; Phu Mu; Phu Hin Khan; Nam Tok Tad Ton)
Graphium doson axion ; Katsuyama et al.,2006 : 46, fig.31,♂. (Nan: Ban Nam Poon)
Graphium doson axion ; Ek-Amnuay,[2007] : 118, pl.31, figs.P65a♂,♂(Un),♀. (Bangkok; Chanthaburi; Chiang Dao)
Graphium doson axion ; Racheli & Cotton,2009 : 44-45, figs.♂,♂(Un). (Chiang Mai)
Graphium doson axion ; Kimura et al.,2011 : 138, figs.♂,♂(Un),♀,♀(Un). (Doi Suthep; Thung Saleng Luang; Bung Kahan; Phu Khiewo; Sarika; Nakohn Nayork; Wan Nam Khiewo; Kao Soi Dao; Khao Khiewo; Doi Musso; Sai York Noi; Khlong Tariew; Khao Phang WF; Ratchaburi; Kaeng Krachan; Hua Hin)
Graphium doson axion ; Ek-Amnuay,2012 : 118, pl.31, figs.P62a♂,♂(Un),♀. (Bangkok; Chanthaburi; Chiang Dao)
Graphium doson axion ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Arisbe doson axionides ; Page & Treadaway,2014 : 272-273. (Chiang Mai)
Arisbe doson actor ; Page & Treadaway,2014 : 273-274, figs.19,♂(Spring form),20,♂(Summer form). (Phrae)
Graphium doson axion ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Sa Kaeo; Chonburi; Chanthaburi) (accessed: 2022.06.01)
Graphium doson axion ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Graphium doson axion ; Motono & Negishi,1989 : 47, pl.9, figs.7,♂,8,♂(Un). (Pakse; Vientiane; TL; Luang Prabang; Nam Ngum Dam)
Graphium doson axion ; Osada, Uémura & Uehara,1999 : 201, pl.26, figs.♂,♂(Un),♂(Un). (Thabok; Lak Sao; Saravan)
Graphium doson axion ; Cotton & Racheli,[2007] : 312-313. (every locality)
Graphium doson axion ; Racheli & Cotton,2009 : 44-45, figs.♀,♀(Un),♂,♂(Un). (Nakai: Ban Nam Nien; Oudomxay)
Graphium doson axion ; Onodera,2009b : 34-35,39-40. (Vientinane: Namsong Riverside; Bolikhamxai: Thabok)
Graphium doson axion ; Nakamura & Wakahara,2012 : 42.
Arisbe doson actor ; Page & Treadaway,2014 : 273-274. (Laos)

[ CAMBODIA ]
Graphium doson axion ; Woodfield & Murton,2006 : figs.♀,♀(Un). (Botum Sakor NP.)
Graphium doson ; Onodera,2007 : 26. (Rattanakiri: Ou Chum)
Graphium doson ; Onodera,2008 : 57. (Mondolkiri: Bousra)
Graphium doson ; Onodera,2009 : 38. (Siem Reap: Phnum Kulen)
Arisbe doson actor ; Page & Treadaway,2014 : 273-274. (Cambodia)

[ VIETNAM ]
Papilio jason axion ; Fruhstorfer,1903 : 204-206. (Tonkin: Than Moi; Chiem-Hoa / Annam: Xom-Gom)
Papilio doson axion ; Jordan,1909 : 97. (Tonkin / Annam)
Papilio doson ; Lemee,1950 : 4. (Backan; Hagiang)
Papilio doson axion f.vern.acheron ; Metaye,1957 : 83. (N / C / S)
Papilio doson axion f.aest.praestabilis ; Metaye,1957 : 83. (N / C / S)
Graphium doson axion ; Ikeda et al.,1998 : 17, figs.4:19,♂,20,♂(Un), 9:14(♂genitalia). (Cuc Phuong)
Graphium doson axion ; Shinkai,1999 : 62. (N: Tam Dao / S: Bao Lac)
Graphium doson ; Vu,2003 : 56. (Phu Quoc)
Graphium doson axion ; Luong et al.,2004 : 1, 27, figs.♂,♂(Un). (Cuc Phuong)
Graphium doson axion ; Miyazaki, Saito & Saito,2006 : 10, fig.Pa-29,♂. (Bin Thuan: Dong Tien; Ho Chi Minh)
Graphium doson ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien)
Graphium doson axion ; Monastyrskii,2007 : 80-81, pl.25, fig.3a♂,3b♂(Un). (throughout Vietnam)
Graphium doson axion ; Miyazaki, Saito & Saito,2007c : 36, fig.Pa-29,♀. (Dong Nai: Thac Mai)
Arisbe doson axionides ; Page & Treadaway,2014 : 272-273. (Ha-Giang)
Arisbe doson actor ; Page & Treadaway,2014 : 273-274. (Dak Lak)
Graphium doson axion ; Monastyrskii & Devyatkin,2015 : 14. (N / C / S)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 12♂ 2♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 4♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Kanchanaburi : 1♂ 3♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 8♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Trat : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Bolikhamxai : 30♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 24♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Champasak : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Ninh Binh : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Dong Nai : 5♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Lam Dong : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is very common in secondary and montane forest at low to moderate elevations (alt.30m - 1300m). Males frequently come to the banks of streams and puddles to suck water.
Page & Treadaway,2014 treated the populations from Nepal to China as two subspecies, ssp.axionides (Pakistan to Hong Kong, N.W.Thailand (Chiang Mai) and 'N.W.'[N.E.]Vietnam (Ha Giang)) and ssp.actor (Guangdong, Hainan, Laos, 'E.'[N].Thailand (Phrae), Cambodia and Vietnam). However, they are quite variable and more subject to seasonal and individual variation than to geographic variation, and cannot be clearly separated.


Literature

Felder, C. & Felder, R.,1864 : Species Lepidopterorum hucusque descriptae vel iconibus expressae in seriem systematicum digestae.
Verh. zool.-bot. Ges. Wien 14:289-378.

Fruhstorfer, H.,1903 : Verzeichnis der in Tonkin, Annam und Siam gesammelten Papilioniden und Besprechung verwandter Formen.
Berl. ent. Z. 47(3/4):167-234.

Fruhstorfer, H.,1907 : Neue und alte Rassen von Papilio jason.
Ent. Zeit. 21(34):209.

Fruhstorfer, H.,1909 : Neue Papilioniden des indo-australischen Faunengebietes.
Ent. Zeit. 22(41):169-170.

Jordan, K.,1909 : Die Indo-Australische Tagfalter. Papilionidae in Seitz, Grossschmett. Erde
9:97-104.

Kocak, A.O. & Kemal, M.,2000: Sergi Asiye ve Pasipiktiki bezi taksonlar heqqide nomenklatur hatiriliri (Papilionidae, Lepidoptera).
Cent. ent. Stud. Misc. 71:1-6, 11.

Jiang, F.Q, Huang, B.K. & Yao, L.H.,2001 : Fauna of insects in Fujian Province of China 4 - Lepidoptera: Rhopalocera.
Fujian Science & Technology, Fuzhou.

Page, M.G.P. & Treadaway, C.G.,2014 : Revisional notes on the Arisbe eurypylus species group (Lepidoptera: Papilionidea: Papilionidae)
Stutt. Beit. Natur. Serie A, Neue Ser. 7:253-284.


Graphium doson actor ♂ Graphium doson actor ♂
♂, Chiang Mai, Thailand. ♂, Chiang Mai, Thailand.
Graphium doson actor ♂
♂, Chiang Mai, Thailand.

1996.10.08 - 2022.06.30.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.