♂, Bolikhamxai, Laos.		♀, Bolikhamxai, Laos.
♂, Bolikhamxai, Laos.		♀, Sekong, Laos.
♂ genitalia, Bolikhamxai, Laos.

♂ (bred), Nakhon Nayok, Thailand.		♂ genitalia, Nakhon Nayok, Thailand.

ON : Thamala marciana crocea Seki & Nakamura, 2026 OD : Butterflies 99: 30-31, figs.6A(HT,♂,♂Un), 6Ag1-5(♂genitalia), 6B(PT,♂,♂Un), 6C(PT,♀,♀Un), 6D(PT,♀,♀Un). TL : Lak 36, near Thakhek, Khammouan, C.Laos. Distribution : C. & E.Thailand, Laos, Cambodia, Vietnam. ● ssp.crocea ● ssp.maia ● ssp.sarupa ● ssp.marciana

Holotype ♂, 28.Mar.2003, Lak 36, near Thakhek, Khammouane, C.Laos, N.Nakamura leg. (in coll. N.Nakamura, will be deposited at RIEB)
Paratypes ('11♂' [12♂] 9♀): Laos: Vientiane, 1♂; Bolikhamxai, 5♂ 4♀; Khammouane, 1♂ 1♀; Attapeu, 1♂ 1♀;
Thailand: 'Phrae: 1♂'; Nakhon Ratchashima: 1♂ 1♀; Vietnam: Lam Dong: 2♂ 2♀. (RIEB ; in coll. Y.Seki ; N.Nakamura)

[ THAILAND ]
Thamala marciana petrella ; Pinratana,1981 : 119, pl.21, fig.15,♂. (Nakhon Ratchasima)
Thamala marciana miniata ; Pinratana,1981 : 119, pl.34, figs.5,♂,6,♀ ; pl.35, fig.3,♀(Un). (Nakhon Ratchasima)
'Thamala marciana marciana ; Pinratana,1981 : 120. not fig. (Chiang Mai)'
Thamala marciana miniata ; Ek-Amnuay,[2007] : 668, pl.305, figs.L321a♂,♂(Un). (Nakhon Ratchasima)
Thamala marciana petrella ; Ek-Amnuay,[2007] : 668, pl.305, figs.L321b♂,♂(Un). (Nakhon Ratchasima)
Thamala marciana miniata ; Ek-Amnuay,2012 : 706, pl.323, figs.L345a♂,♂(Un),♂,♂(Un). (Nakhon Ratchasima)
Thamala marciana miniata ; Kimura et al.,2014 : 101, figs.♂,♂(Un),♀,♀(Un). (Wan Nam Khiewo)
Thamala marciana petrella ; Kimura et al.,2014 : 101. (by literature)
Thamala marciana ssp. ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2023- : Butterflies of Thailand, figs., Early Stages. (Ubon Ratchathani; Sisaket; Sa Kaeo; Nakhon Ratchasima; Nakhon Nayok; Prachinburi) (accessed: 2024.06.08)
Thamala marciana subsp. incertae sedis ; Inayoshi,2019-2024 : Butterflies in Indo-China, figs.1♂,(genitalia). (Nakhon Nayok) (30.Jun.2024 last updated ; 30.Jun.2026 changed.)

[ LAOS ]
Thamala marciana miniata ; Motono & Negishi,1989 : 75, pl.107, figs.A3,♀,A4,♀(Un),A5,♂(Un). (Tha Ngone; Nam Moh)
Thamala marciana miniata ; Osada, Uémura & Uehara,1999 : 218, pl.123, figs.♂,♀. (Lak Sao)
Thamala marciana miniata ; Nimura & Wakahara,2005 : 48. (Namdhua; Thabeng)
Thamala marciana miniata ; Onodera,2009b : 43. (Bolikhamxai: Phon Si)
Thamala marciana miniata ; Nakamura & Wakahara,2012 : 54.
Thamala marciana miniata ; Onodera,2022 : 94, pl.227, figs.♂,♂(Un),♀,♀(Un). (Khammouan: Nahin; Bolikhamxai: Phon Si)
Thamala marciana subsp. incertae sedis ; Inayoshi,2019-2024 : Butterflies in Indo-China, figs.2♂2♀, 1♂(genitalia). (Bolikhamxai; Sekong) (30.Jun.2024 last updated ; 30.Jun.2026 changed.)

[ CAMBODIA ]
Thamala marciana ; Onodera,2009 : 41, figs.♂[♀],♂[♀](Un). (Siem Reap: Phnum Kulen)
Thamala marciana miniata ; Onodera,2022 : 129, pl.267, figs.♀,♀(Un). (Siem Reap: Phnum Kulen)
Thamala marciana ; Chartier & Kosterin,2022 : 123. (Siem Reap)

[ VIETNAM ]
Thamala marciana ; Monastyrskii et al.,2006 : 43. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Thamala marciana miniata ; Miyazaki, Saito & Saito,2006d : 64, figs.L-111,♂,♀. (Lam Dong: Dambri)
Thamala marciana miniata ; Monastyrskii & Devyatkin,2015 : 64. (C / S)

[ THAILAND ]
Nakhon Nayok : 1♂ 1♀ (photo by M.Hayashi and S.Sophonviwatkul) (1♂ genitalia dissected).

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[ LAOS ]
Bolikhamxai : 3♂ 1♀ (1♂ genitalia dissected).

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Sekong : 1♀.

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The butterfly is uncommon in montane forests at low to moderate elevations.
The populations from C. Thailand, Laos, and Vietnam resemble ssp.miniata from Upper Tenasserim, Myanmar. However, females of ssp.miniata have prominent whitish scales in the tornal area on the UpH, whereas those from the former populations virtually lack them. Due to significant individual and geographical variation within the Myanmar populations, a comprehensive review of the entire mainland population appears necessary.
Seki & Nakamura (2026) recorded this taxon from "Wan Chin, Phrae, N. Thailand", but this record is considered dubious. To date, I have not personally examined or confirmed any specimens of this species from N. Thailand. Consequently, the record from Phrae is excluded from the distribution map pending the availability of verifiable collection records.
Interestingly, the larval maculation in C. Thailand (Nakhon Nayok, and presumably also Nakhon Ratchasima) differs significantly from that in Laos and Peninsular Thailand. However, the wing markings and the male genitalia are remarkably similar to those of the population from Laos. It is possible that this population differs slightly in its molecular characters from the populations spanning E. Thailand to Vietnam. This population is provisionally included in this taxon herein, however, further taxonomic studies, particularly incorporating molecular analysis, are warranted.

The difference between the final instar larvae of Nakhon Nayok and Yala.
'Thamala marciana crocea' Nakhon Nayok, Thailand. (photo by S.Sophonviwatkul)	Thamala marciana marciana Yala, Thailand. (photo by S.Atdhabhan)

Seki, Y. & Nakamura, N.,2026 : New findings on Lycaenidae (Lepidoptera) from Indochina, including descriptions of a new species and three new subspecies.
Butterflies 99: 25-42.

Seitz, A.,1926 : Family Lycaenidae, In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde.
9: 986, pls. (part), Alfred Kernen, Stuttgart.

Corbet, A.S.,1942 : Miscellaneous notes on the butterflies of the Malay Archipelago.
Entomologist 75: 172-174.

Corbet, A.S.,1944 : Some notes on the Butterflies of the Langkawi Islands.
Entomologist 77(3): 39-42.

Igarashi, S. & Fukuda, H.,2000 : The Life Histories of Asian Butterflies vol.2
Tokai University Press, Tokyo. 711 pp, 427 col.pls.

♀, Nakhon Nayok, Thailand. (photo by M.Hayashi) 2024.06.30. A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.