♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Luang Phabang, Laos. (photo by H.Onodera)		♀, Khammouan, Laos. (photo by H.Onodera)
♂, Nan, Thailand.		♂, Chiang Mai, Thailand.

ON : Flos yunnanensis Wang & Fan, 2002 OD : Butterflies Fauna Sinica: Lycaenidae. 143, pl.11, fig.19,♀,20,♀(Un) as Flos gunnanensis. TL : Mengala county, Yunnan. (♀) Distribution : N.Thailand, Laos, Vietnam, Yunnan. ● ssp.yunnanensis ● ssp.artegal ● ssp.anniella

Flos gunnanensis Wang & Fan, 2002 : Butterflies Fauna Sinica: Lycaenidae. pl.11, figs.19,♀,20,♀(Un). (IOS)

Holotype ♀, 12.Apt.1994. Mengla County, Yunnan. Wang & Fan leg. (Entomological Museum, Northwestern Agricultural University)

[ THAILAND ]
Flos anniella artegal ; Pinratana,1981 : 104. (Chiang Mai) (in part)
Flos anniella artegal ; Ek-Amnuay,[2007] : 650. (Chiang Mai) (in part)
Flos anniella artegal ; Ek-Amnuay,2012 : 686. (Chiang Mai) (in part)
Flos anniella artegal ; Kimura et al.,2014 : 85, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Doi Phui; Doi Suthep; Nam Tok Mae Sa)
Flos anniella yunnanensis ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2022- : Butterflies of Thailand, fig, Early Stages. (Chiang Mai) (accessed: 2022.06.16)

[ LAOS ]
Flos anniella artegal ; Osada, Uémura & Uehara,1999 : 217, pl.119, figs.♂,♂(Un),♀. (Lak Sao; Xaignabouri)
Flos anniella artegal ; Nimura & Wakahara,2005 : 47. (Nakai)
Flos anniella artegal ; Nakamura & Wakahara,2012 : 54.

[ VIETNAM ]
Flos anniella ; Monastyrskii et al.,2006 : 43. (Thua Thien Hue: Nam Dong)
Flos anniella artegal ; Miyazaki, Saito & Saito,[2007] : 61, figs.L-97,♂,♀. (Lam Dong: Dambri; Dai Duc Me)
Flos anniella artegal ; Monastyrskii & Devyatkin,2015 : 62. (C: Nghe An to Lam Dong)

[ THAILAND ]
Mae Hong Son : 1♂ 1♀.

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Chiang Mai : 10♂ 5♀ (incl.photo).

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Nan : 1♂.

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Kamphaeng Phet : 1♀ (photo by R.Charoenpon).

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[ LAOS ]
Luang Phabang : 2♂ (photo by H.Onodera).

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Xieng Khouang : 1♂ (photo by H.Onodera).

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Khammouan : 1♀ (photo by H.Onodera).

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The butterfly is common in montane forest at low to high elevations (Alt.300m - 1600m). Males are territorial on trees or shrubs at hilltops or forest edges.
Wang & Fan,2002 described Flos yunnanensis as a distinct species, but it's considered to be Flos anniella by characteristic markings of the underside.
On the other hand, the populations of anniella from N.Thailand, Laos and Vietnam were hitherto treated as ssp.artegal, However female of this populations have very broad borders on both wings same as yunnanensis. Male has darker ground color and more inconspicuous markings on the UnH. than other subspecies. Therefore, I treat Flos yunnanensis to subspecies of Flos anniella and including Northern Indochinese taxon to this subspecies.
It is very interesting whether the populations of W.Thailand (Tak & Kanchanaburi) are this subspecies or ssp.artegal. Further taxonomic study and distributional research are needed by female specimens.

Wang, M. & Fan, X.,2002 : Butterflies Fauna Sinica: Lycaenidae.
Henan Science and Technology Publishing House. 439p., 28 pls.

♂, Chiang Mai, Thailand.	♀, Mae Hong Son, Thailand.
♂ (territorial behavior), Chiang Mai, Thailand.
♂ (territorial behavior), Chiang Mai, Thailand.	♀, Kamphaeng Phet, Thailand. (photo by R.Charoenpon)