♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♂, Luang Phabang, Laos. (photo by H.Onodera)		♀, Khammouan, Laos. (photo by H.Onodera)
♂, Nan, Thailand.		♂, Chiang Mai, Thailand.

ON : Flos yunnanensis Wang & Fan, 2002 OD : Butterflies Fauna Sinica, Lycaenidae: 143, pl.11, fig.19,♀, 20,♀(Un) [as Flos gunnanensis]. TL : Mengala county, Yunnan. Distribution : Yunnan, N.Thailand, Laos, Vietnam. ● ssp.yunnanensis ● ssp.artegal ● ssp.anniella

Flos gunnanensis Wang & Fan, 2002 : Butterflies Fauna Sinica, Lycaenidae: pl.11, figs.19,♀, 20,♀(Un). (IOS)

Holotype ♀, 12.Apt.1994. Mengla County, Yunnan. Wang & Fan leg. (Entomological Museum, Northwestern Agricultural University)

[ THAILAND ]
Flos anniella artegal ; Pinratana,1981 : 104. (Chiang Mai) (in part)
Flos anniella artegal ; Ek-Amnuay,[2007] : 650. (Chiang Mai) (in part)
Flos anniella artegal ; Ek-Amnuay,2012 : 686. (Chiang Mai) (in part)
Flos anniella artegal ; Kimura et al.,2014 : 85, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Doi Phui; Doi Suthep; Nam Tok Mae Sa)
Flos anniella yunnanensis ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2022- : Butterflies of Thailand, figs., Early Stages. (Chiang Mai) (accessed: 2022.06.16)

[ LAOS ]
Flos anniella artegal ; Osada, Uémura & Uehara,1999 : 217, pl.119, figs.♂,♂(Un),♀. (Lak Sao; Xaignabouri)
Flos anniella artegal ; Nimura & Wakahara,2005 : 47. (Nakai)
Flos anniella artegal ; Nakamura & Wakahara,2012 : 54.
Flos anniella artegal ; Onodera,2022 : 88, pl.219, figs.♂,♂(Un),♀,♀(Un). (Luang Phabang: Marphouk; Khammouan: Beungna)

[ VIETNAM ]
Flos anniella ; Monastyrskii et al.,2006 : 43. (Thua Thien Hue: Nam Dong)
Flos anniella artegal ; Miyazaki, Saito & Saito,2006d : 61, figs.L-97,♂,♀. (Lam Dong: Dambri; Dai Duc Me)
Flos anniella artegal ; Monastyrskii & Devyatkin,2015 : 62. (C: Nghe An to Lam Dong)

[ THAILAND ]
Mae Hong Son : 1♂ 1♀.

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Chiang Mai : 10♂ 5♀ (incl. photo).

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Nan : 1♂.

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[ LAOS ]
Luang Phabang : 2♂ (photo by H.Onodera).

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Xieng Khouang : 1♂ (photo by H.Onodera).

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Khammouan : 1♀ (photo by H.Onodera).

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The butterfly is common in montane forests at low to high elevations (Alt. 300 - 1750m). Males are territorial on trees or shrubs at hilltops.
Wang & Fan (2002) described Flos yunnanensis as a distinct species, but the distinctive markings on the UnH suggests that it is Flos anniella. On the other hand, the populations of anniella from N. Thailand, Laos and Vietnam have been treated as ssp.artegal. However, the females of these populations have very broad black margins on both wings, the same as yunnanensis. The males have a darker ground color and less distinctive markings on the UnH than other subspecies. Therefore, I treat Flos yunnanensis as a subspecies of Flos anniella and include the northern Indochinese populations in this taxon.

Wang, M. & Fan, X.,2002 : Butterflies Fauna Sinica: Lycaenidae.
Henan Science and Technology Publishing House. 439 pp., 28 pls.

♂, Chiang Mai, Thailand.	♀, Mae Hong Son, Thailand.
♂ (territorial behavior), Chiang Mai, Thailand.
♀, Chiang Mai, Thailand.
♂ (territorial behavior), Chiang Mai, Thailand.	♂ (territorial behavior), Chiang Mai, Thailand.

2019.06.30 - 2024.01.31.