♂, Bangkok, Thailand.		♀, Mae Hong Son, Thailand.
♂, Tak, Thailand.		♀, Bolikhamxai, Laos.
♂, Yala, Thailand.		♀, Ranong, Thailand.
♂, Satun, Thailand.		♂, Surat Thani (Koh Phangan dist.), Thailand.

ON : Papilio otis Fabricius, 1787 OD : Mantissa Ins. 2:73-74. TL : China. Distribution : Andaman, Myanmar (Kachin to Shan State), Thailand, Laos, Cambodia, Vietnam, Sichuan, Guangxi, Hainan, Langkawi, W.Malaysia, Tioman, Aur, Singapore, Borneo, Philippines, Guam, Palau, Moen Is., Bangka,

Polyommauts sangra Moore, [1866] : Proc. zool. Soc. Lond. 1865(3):772, pl.41, fig.8. TL. Bengal.
Zizera oriens Butler, 1883 : Ann. Mag. nat. Hist. (5)11(66):417-418. TL. Pasananca valley, Mindanao.
Zizeeria otis lampa Corbet, 1940 : Entomologist 83:41. TL. Singapore ; Malacca.

lampa ; Type, ♂, Jan.1930. Singapore. W.H.Evans ; Type, ♀, Malacca.

[ THAILAND ]
Zizina otis ; Godfrey,1916 : 135.
Zizina otis ; Pendlebury,1923 : 44. (Nakhon Sri Thammarat: Tai Sai River; Wat Kiriwong)
Zizera otis ; Godfrey,1930 : 319. (Thailand)
Zizina otis ; Godfrey,1932a : 271. (P)
Zizina otis otis ; Shirôzu & Saigusa,1962 : 85-87, pl.17, figs.127,♂, 128,♂(Un), Textfig.44,♂genitalia. (Bangkok; Hin Kang; Nakon Nayok; Li; Kasetsar; Chieng Mai; Doi Inthanon; Pon Makampon)
Zizina otis otis ; Kawazoé,1964 : 152. (N: Ban Tak; P: Kaper)
Zizina otis sangra ; Pinratana,1981 : 50, pl.7, figs.26,♂,27,♂(Un). (Nakhon Nayok; Ratchaburi)
Zizina otis lampa ; Pinratana,1981 : 50, pl.7. not fig. (P)
Zizina otis sangra ; Yamaguchi & Aoki,2002 : 34, fig.96,♂. (Mukdahan: Ban Lao Kram)
Zizina otis sangra ; Katsuyama et al.,2005 : 66, fig.72,♂. (Umphang: Tha Sai; Umphang)
Zizina otis sangra ; Katsuyama et al.,2006 : 47, fig.51,♂. (Nan: Ban Nam Poon)
Zizina otis sangra ; Ek-Amnuay,[2007] : 568, pl.257, figs.L99a♂,♂(Un),♂,♀,♀(Un),♀. (Bangkok)
Zizina otis lampa ; Ek-Amnuay,[2007] : 568, pl.257, figs.L99b♀,♀(Un). (Song Khla; Khao Phanon Bencha)
Zizina otis lampa ; Les Day,2011-2022 : Butterflies of Koh Samui, figs.♂,♀,[♂](Un). (Koh Samui) (accessed: 2021.01.30)
Zizina otis sangra ; Ek-Amnuay,2012 : 588, pl.266, figs.L124a♂,♂(Un),♀. (Bangkok)
Zizina otis lampa ; Ek-Amnuay,2012 : 588, pl.266, figs.L124b♂,♀(Un),♀. (Songkhla; Khao Phanon Bencha)
Zizina otis sangra ; Kimura et al.,2014 : 180, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Ang Khang; Chiang Dao; Doi Inthanon; Khun Lao Village; Phu Khiewo; Thoen; Nakohn Nayok; Bang Phra; Bangkok; Doi Musso; Kaeng Chai; Bang Pain; Kanchanaburi; Wang Keo)
Zizina otis lampa ; Kimura et al.,2014 : 180, figs.♂,♂(Un),♀,♀(Un). (Ban Tung Tong; Sonkhala; Than To WF.)
Zizina otis sangra ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Chiang Mai; Nakhon Nayok; Bangkok) (accessed: 2023.03.26)
Zizina otis sangra ; Chaianunporn & Chaianunporn,2019 : 85. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Zizina otis sangra ; Motono & Negishi,1989 : 70, pl.100, figs.B5,♂,B6,♂(Un),B7,♀,B8,♀(Un). (Vientiane; TT; Luang Prabang; Phatong)
Zizina otis sangra ; Osada, Uémura & Uehara,1999 : 215, pl.108, figs.♂(dsf),♀(dsf)♂,♂(Un). (Lak Sao)
Zizina otis sangra ; Nimura & Wakahara,2005 : 46. (Namdhua; Thakhek)
Zizina otis sangra ; Nakamura & Wakahara,2012 : 51.
Zizina otis sangra ; Onodera,2022 : 74, pl.206, figs.♂,♂(Un),♀,♀(Un). (Vientiane: Vang Vieng; Khammouan: Natung)

[ CAMBODIA ]
Zizina otis ; Onodera,2007 : 22, fig.80,♂. (Rattanakiri: Poy)
Zizina otis ; Onodera,2008 : 56. (Mondolkiri: Leu)
Zizina otis sangara ; Monastyrskii et al.,2011 : 130. (Phnom Samkos W.S.)
Zizina otis sangara ; Chartier & Kosterin,2022 : 121. (Koh Kong; Mondulkiri; Phnom Penh; Preah Sihanouk; Pursat; Preah Vihear; Ratanakiri; Siem Reap)

[ VIETNAM ]
Zizina otis otis ; Metaye,1957 : 90. (no loc.)
Zizina otis indica ; Metaye,1957 : 90. (no loc.)
Zizina otis sangra ; Ikeda et al.,2001 : 48, figs.2:15,♂,16,♂(Un),6:14(♂genitalia). (Cuc Phuong)
Zizina otis ; Vu,2003 : 58. (Phu Quoc)
Zizina otis sangara ; Luong et al.,2004 : 9, 75, figs.♂,♂(Un). (Cuc Phuong)
Zizina otis ; Monastyrskii et al.,2006 : 42. (Thua Thien Hue: Huong Thuy)
Zizina otis sangra ; Miyazaki, Saito & Saito,[2007] : 52, figs.L-19,♂,♀. (Lam Dong: Dambri; Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien; Ho Chi Minh; Phu Quoc)
Zizina otis sangra ; Nihira,2009 : 100. (Phu Quoc)
Zizina otis sangra ; Monastyrskii & Devyatkin,2015 : 51. (N / C / S)
Zizina otis sangara ; Saito et al.,2017 ; 80. (S: Nui Cam; Nui Coto; Nui Ket)

[ THAILAND ]
Mae Hong Son : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 6♂ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Bangkok : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Trat : 2♂ (photo by E.Ikari).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Ranong : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Surat Thani (Koh Tao ; Koh Phangan) : 6♂ 1♀ (incl. photo by P.Kosierb, A.Giudici).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Satun : 3♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Yala : 4♂ 1♀ (incl. photo by S.Atdhabhan).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Bolikhamxai : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Champasak : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is locally common in fields, open areas, wasteland and roadsides around secondary forests at low to moderate elevations (Alt. 50 - 1200m). Both sexes visit flowers.

Fabricius, J.C.,1787 : Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis,
differentiis specificis, emendationibus, observationibus.
Copenhagen, Christ. Gottl. Proft 8vo 1:xx+348pp ; 2:[2]+382pp

Moore, F.,[1866] : On the lepidopterous insects of Bengal.
Proc. zool. Soc. Lond. 1865(3):755-822, 3 pls.

Butler, A.G.,1883 : The Lepidoptera collected during the recent expedition of H.M.S. "Challenger."
Ann. Mag. nat. Hist. (5)11(66):402-428.

Corbet, A.S.,1940 : Revisional notes on Malayan Rhopalocera.
Entomologist 73:39-43.

Yago, M., Hirai, N., Kondo, M., Tanikawa, T., Ishii, M., Wang, M., Williams, M. & Ueshima, R.,2008 :
Molecular systematics and biogeography of the genus Zizina (Lepidoptera: Lycaenidae)
Zootaxa 1746:15-38.

♂, Chiang Mai, Thailand.
♂, ♀+♂ (mating), Surat Thani (Koh Phangan dist.), Thailand. (photo by A.Giudici)

1999.07.04 - 2023.04.30.