♂, Trat, Thailand.		♀, Trat, Thailand.
♂, Trat, Thailand.		♀, Trat, Thailand.
♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.
♀, Chanthaburi, Thailand.		♂ genitalia, Chiang Mai, Thailand.

ON : Neptis camboja Moore, 1879 OD : Proc. zool. Soc. Lond. 1879(1):136. TL : Cambodia. Distribution : India (Assam), Myanmar, Thailand, Laos, Cambodia, Vietnam, Langkawi.

Rahinda sattanga Moore, 1881 : Trans. ent. Soc. Lond. 29(3):311. TL. "British Burmah" [Myanmar]. (NHML)
Neptis kuhasa Nicéville, 1886 : Butts. India Burmah Ceylon. 2:84. TL. Irangmara, Cachar, [Assam, India]. (NZC).

kuhasa ; Syntypes 1♂, 29.Jul.1881. Cachar, Assam. (NZC), 1♀, 8.Aug.1881. Cachar, Assam (NZC) ; Sheela et al.,2019 : 221, figs.♂(Un),♂, 222, figs.♀,♀(Un).

[ THAILAND ]
Lasippa tiga camboja ; Eliot,1969 : 43. (Siam)
Lasippa tiga camboja ; Yamaguchi & Aoki,2002 : 28, fig.75,♂. (Mukdahan: Phu Huay Sing)
Pantoporia sandaka davidsoni ; Ek-Amnuay,[2007] : pl.174, figs.N97,♂,♂(Un). (in part)
Lasippa camboja ; Ek-Amnuay,[2007] : 398, pl.173, fig.N92,♀. (in part)
Pantoporia sandaka davidsoni ; Ek-Amnuay,2012 : pl.170, figs.N230,♂(left),♂(left,Un). (in part)
Lasippa tiga camboja ; Ek-Amnuay,2012 : 396, pl.171, fig.N236a♀. (in part)
Lasippa camboja ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Sa Kaeo; Nakhon Ratchasima; Phetchaburi) (accessed: 2024.05.12)
Lasippa tiga camboja ; Kimura et al.,2016 : 70-71, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Phu Khiewo; Wan Nam Khiewo; Kao Soi Dao; Kao Khiewo; Kaper) (in part)

[ LAOS ]
Lasippa tiga ssp. ; Osada, Uémura & Uehara,1999 : 206, pl.57, figs.♂,♂(Un). (Vang Vieng; Lak Sao)
Lasippa tiga ssp. ; Nimura & Wakahara,2005 : 42. (Namdhua)
Lasippa tiga camboja ; Nakamura & Wakahara,2012 : 45.
Lasippa tiga camboja ; Onodera,2022 : 37, pl.114, figs.♂,♂(Un),♀,♀(Un). (Bolikhamxai: Phon Si; Champasak: Dong Huasao)

[ CAMBODIA ]
Neptis cambodga [sic] ; Vitalis,1919 : 229. (Cambodge)
Neptis sattanga ; Vitalis,1919 : 229. (Cambodge)
Lasippa tiga siaka ; Kosterin,2019 : 97, fig.49. (Preah Sihanouk)
Lasippa camboja ; Chartier & Kosterin,2022 : 117. (Koh Kong; Mondulkiri; Preah Sihanouk; Siem Reap)

[ VIETNAM ]
Neptis sattanga ; Vitalis,1919 : 229. (Cochinchine)
Lasippa tiga ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong)
Lasippa tiga camboja ; Miyazaki, Saito & Saito,2006b : 22, figs.N-61,♂,♀. (Lam Dong: Dambri)
Lasippa tiga camboja ; Nihira,2009 : 99, fig.♂[♀]. (Phu Quoc)
Lasippa heliodore heliodore ; Yokochi et al.,2015 : 40, figs.55,♂,55v♂(Un),56,♀,56v♀(Un). (Khanh Hoa: Hon Ba)
Lasippa tiga camboja ; Monastyrskii & Devyatkin,2015 : 36. (C / S)
Lasippa heliodore heliodore ; Monastyrskii,2019 : 4(2):52-53, pl.42, figs.1a♂,1b♂(Un),1c♀,1d♀(Un). (C: Pu Mat; Phong Dien; Huong Son; Phong Nha Ke Bang; Bach Ma; A Luoi; Nam Dong; Kon Cha Rang; Chu Yang Sin; Cat Loc; Da Teh; Loc Bac; Bao Lam / S: Cat Tien; Lo Go Xa Mat; La Nga; Nghia Trung)
Lasippa tiga camboja ; Monastyrskii,2019 : 4(2):53-54, pl.42, figs.2a♀,2b♀(Un). (N: Ba Be; Cat Ba; Cuc Phuong / C: Pu Mat; Phong Nha Ke Bang; Phong Dien; Ben En; Vu Quang; Hoang Son; Nam Dong; Loc Bao; Bao Lam; Dong Xuan / S: Cat Tien)

[ THAILAND ]
Mae Hong Son : 2♂.

Jan

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Dec

Chiang Mai : 4♂ 1♀ (incl. photo).

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Chanthaburi : 1♀.

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Trat : 4♂ 4♀ (incl. photo).

Jan

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Phetchaburi : 1♂ (photo by T.Aoyama).

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[ LAOS ]
Bolikhamxai : 1♂ (photo by H.Onodera).

Jan

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[ VIETNAM ]
Lam Dong : 1♂ 3♀,

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The butterfly is locally common in montane forest at low to moderate elevations (Alt. 20 - 700m). Males are territorial on trees about 2 to 3m high at forest edges and are very cautious.
This taxon and Lasippa tiga siaka are found sympatrically in peninsular to S.E. Thailand, furthermore, the male genitalia is different. Valva of camboja is thin and longer than tiga and heliodore. Therefore I consider camboja ia a distinct species as Eliot (1969) and Kimura et al. (2016) pointed out.

Moore, F.,1879 : Descriptions of new Asiatic diurnal Lepidoptera.
Proc. zool. Soc. Lond. 1879(1):136-144.

Moore, F.,1881 : Descriptions of new Asiatic diurnal Lepidoptera.
Trans. ent. Soc. Lond. 29(3):305-313.

Nicéville, L. de,1886 : The butterflies of India, Burmah and Ceylon.
A descriptive handbook of all the known species of rhopalocerous Lepidoptera inhabiting that region, with notices of allied species occurring in the neighbouring countries along the border, with numerous illustrations. Volume II (Nymphalidae, Libytheidae, Nemeobiidae).
Calcutta Central Press : vii + 332 pp, frontispiece, pls.18-24.

Eliot, J.N.,1969 : An Analysis of the Eurasian and Australian Neptini (Lepidoptera: Nymphalidae).
Bull. Br. Mus. nat. Hist. (Ent.) Supple.15. London.

♂, Trat, Thailand.
♂, Chiang Mai, Thailand.	♂ (territorial behavior), Chiang Mai, Thailand.
♂, Phetchaburi, Thailand. (photo by T.Aoyama)

2006.02.20 - 2024.05.31.