♂, Champasak, Laos.		♂, Bolikhamxai, Laos. (photo by H.Onodera)
♂, Lam Dong, Vietnam. (photo by K.Saito)		♂, Lam Dong, Vietnam. (photo by K.Saito)
♂ genitalia, Champasak, Laos.		♂ genitalia, Lam Dong, Vietnam. (photo by K.Saito)

ON : Isma dawna minuscla Inoué & Kawazoé, 1967 OD : Trans. lepid. Soc. Japan 17(1/2):9-10, figs.123(HT,♂), 124(HT,♂Un), tf.35(♂,genitalia). TL : Trang Bom, S.Vietnam. Distribution : Laos, S.Vietnam. ● ssp.minuscla ● ssp.umbrosa

Isma umbrosa contracta Inoué & Kawazoé, 1967 : Trans. lepid. Soc. Japan 17(1/2):10-11, figs.125(HT,♂), 126(HT,♂Un), tf.36(♂,genitalia). TL. Trang Bom, S.Vietnam.

minuscla ; Holotype ♂, 18.Dec.1969.Dec. Trang Bom, S.Vietnam.
contracta ; Holotype ♂, 18.Feb.1962. Trang Bom, S.Vietnam.

[ LAOS ]
Isma umbrosa minuscla ; Masui & Uehara,1996 : 5, pl.2, figs.26,♂,♂(Un),♀,♀(Un). (Saravan)
Isma umbrosa minuscla ; Osada, Uémura & Uehara,1999 : 224, pl.140, figs.♂,♀. (Phong Si; Thateng)
Isma minuscla ; Inayoshi,1999 - 2023 : Butterflies in Indo-China, figs.♂,♂(Un),♂,♂(Un),♂(genitalia). (Champasak; Bolikhamxai) (31.Dec.2017 last updated ; 31.Aug.2023 changed.)
Isma umbrosa minuscla ; Nakamura & Wakahara,2012 : 59.
Isma umbrosa minuscla ; Onodera,2022 : 118, pl.253, figs.♂,♂(Un). (Khammouan: Nahin)

[ VIETNAM ]
Isma umbrosa ; Devyatkin & Monastyrskii,2002 : 149. (Khe Lau; Khe Ba Long; Huong Son)
Isma umbrosa ; Monastyrskii et al.,2006 : 45. (Thua Thien Hue: Nam Dong; Phong Dien)
Isma umbrosa minuscla ; Miyazaki et al.,2010 : 29, fig.H-149,♂. (Lam Dong: Gung Re)
Isma dawna minuscla ; Monastyrskii & Devyatkin,2015 : 80. (S: Dong Nai)
Isma umbrosa contracta ; Monastyrskii & Devyatkin,2015 : 80. (C / S)
Isma umbrosa contracta ; Inayoshi,2017 - 2023 : Butterflies in Indo-China, figs.♂,♂(Un),♂,♂(Un),♂(genitalia). (Lam Dong) (31.Dec.2017 last updated ; 31.Aug.2022 changed.)

[ LAOS ]
Bolikhamxai : 1♂ (photo by H.Onodera).

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Champasak : 1♂.

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[ VIETNAM ]
Lam Dong : 2♂.

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The butterfly is uncommon in montane forests at low to moderate elevations (Alt. 150 - 650m).
Inoué & Kawazoé (1967) described this taxon as a subspecies of Isma dawna. Subsequently, Eliot (1973:13) treated both Isma dawna minuscla and Isma umbrosa contracta as synonyms of umbrosa, and he chose minuscla for the subspecies name of Vietnamese umbrosa. The markings of the populations from Laos and Vietnam appear to be rather variable and I agree with Eliot's treatment that both taxa are conspecific. However, the shape of the valva is rather different in umbrosa from peninsular Thailand. Further taxonomic study is needed.

Inoué, S. & Kawazoé, A.,1967 : Hesperiid butterflies from South Vietnam. (4)
Trans. lepid. Soc. Japan 17(1/2):1-17, 128 figs.

Eliot, J.N.,1973 : More revisional Notes on oriental Butterflies.
Entomologist 106(1316):1-15, 10figs.

2017.07.31 - 2023.08.31.