♂, Nakhon Nayok, Thailand. (photo by S.Sophonviwatkul)		♂, Phetchabun, Thailand. (photo by A.Giudici)

ON : Isma vulso bicolor Evans, 1926 OD : J. Bombay nat. Hist. Soc. 31:443. TL : Dawnas. (♂, NHML) Distribution : Myanmar, Thailand, Laos, Cambodia. ● ssp.bicolor ● ssp.obscura

[ THAILAND ]
Isma protoclea ; Godfrey,1916 : 144. (Chonburi: Hup Bon)
Isma protoclea ; Godfrey,1930 : 375. (Chonburi: Hup Bon)
Isma obscura bicolor ; Pinratana,1985 : 75, pl.25, fig.157a♀. (Nakhon Nayok; Chanthaburi)
Isma protoclea bicolor ; Kimura et al.,2011 : 76, figs.♂,♂(Un). (Kao Yai; Kao Soi Dao; Kratching WF.)
Isma protoclea bicolor ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Phetchabun; Nakhon Nayok; Chanthaburi) (accessed: 2023.08.04)

[ LAOS ]
Isma iapis iapis ; Nakamura et al.,2009 : 25, fig.8,♂,♂(Un). (Bolikhamxai: Nam Kata; Nahin; Phongxi)
Isma iapis iapis ; Nakamura & Wakahara,2012 : 59, pl.25, figs.He-145,♂,♂(Un). (Bolikhamxai: Namkata)

[ CAMBODIA ]
Isma protoclea bicolor ; Chartier & Kosterin,2022 : 125, fig.7M,[♂(Un)]. (Koh Kong)

[ THAILAND ]
Phetchabun : 1♂.

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Nakhon Nayok : 1♂.

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The butterfly is uncommon in montane forests at low to moderate elevations (Alt. 150 - 800m).
The record from Laos appears to be this taxon rather than Isma iapis. The populations from the continental region of Indochina have been treated as ssp.bicolor. However, they may be separate taxa from a biogeographical point of view. Further taxonomic and distributional studies are needed.

Evans, W.H.,1926 : The Identification of Indian Butterflies (9-11)
J. Bombay nat. Hist. Soc. 31(1):49-83, pl.30 ; (2):427-446, pl.31 ; (3):615-637, pl.32.

♂, Nakhon Nayok, Thailand. (photo by S.Sophonviwatkul)

1999.09.11 - 2023.08.31.