Tamela nigrita diocles  (Moore, [1866])

ON : Nisoniades diocles Moore, [1866]
OD : Proc. zool. Soc. Lond. 1865(3):787.
TL : Bengal.
Distribution : Nepal, Sikkim to Myanmar,
Vietnam, Guangxi, Hainan.
● ssp.maura
Tamela nigrita diocles map

Records taken from Literature

Kerana diocles ; Lemée,1950 : 25. (Hagiang; Backan)
Ancistroides nigrita diocles ; Devyatkin & Monastyrskii,1999 : 168. (Tam Dao; Ba Be; Na Hang)
Ancistroides nigrita diocles ; Ikeda et al.,2001b : 63, figs.3:9,♂,10,♂(Un),11,♀,5:25(♂genitalia). (Cuc Phuong)
Ancistroides nigrita diocles ; Luong et al.,2004 : 14, 88, figs.♂,♂(Un). (Cuc Phuong)
Ancistroides nigrita ; Monastyrskii et al.,2006 : 45. (Thua Thien Hue: A Luoi)
Ancistroides nigrita diocles ; Monastyrskii & Devyatkin,2015 : 79. (N / C)


Zhang et al. (2022:39) suggested that the taxon should be treated in genus Tamela based on molecular analysis. Subsequently, Zhang et al. (2023:17) treated both taxa diocles and maura as distinct species based on molecular analysis. However, important populations in the Indochina region have not been examined. I treat this as conventionally until this is resolved. Further taxonomic study is needed.


Moore, F.,[1866] : On the lepidopterous insects of Bengal.
Proc. zool. Soc. Lond. 1865(3):755-822, 3 pls.

Zhang, J., Cong, Q., Shen, J. & Grishin, N.V.,2022 : Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera)
Insecta Mundi 0921:1-135.

Xue, G., Qu, P., Li, M., Chiba, H. & Li, W.,2022 :
Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae)
Zootaxa 5205(5):481–490.

Zhang, J., Cong, Q., Shen, J., Song, L., Opler, P.A. & Grishin, N.V.,2023 :
Additional taxonomic refinements suggested by genomic analysis of butterflies
The Taxonomic Report of the International Lepidoptera Survey 11(1):1-25.

1999.09.04 - 2023.09.30.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.