♂, Nan, Thailand.		♀, Mae Hong Son, Thailand.
♂, Nan, Thailand.		♀, Chiang Mai, Thailand.
♂, Yala, Thailand.		♀, Ranong, Thailand.

ON : Nisoniades salsala Moore, 1866 OD : Proc. zool. Soc. Lond. 1865(3):786-787. TL : Bengal. (♂, NHML) Distribution : Kumaon to Myanmra, Thailand, Laos, Cambodia, Vietnam, Hainan, Hong Kong, China, Langkawi, W.Malaysia, Tioman, Singapore, Sumatra, Java.

Iambrix salsala vasuba Fruhstorfer, 1910 : Deut. ent. Zeit. [Iris] 24:93. TL. Montes-Manson, Tonkin. (♂, NHML)

[ THAILAND ]
Iambrix salsala ; Godfrey,1916 : 144.
Iambrix salsala ; Pendlebury,1923 : 48. (Nakhon Sri Thammarat: Tai Sai River; Khao Ram; Lan Saka)
Iambrix salsala salsala ; Godfrey,1927 : 127. (N)
Iambrix salsala salsala ; Godfrey,1930 : 368-369. (except P)
Iambrix salsala salsala ; Godfrey,1931 : 196. (Ko Tao)
Iambrix salsala salsala ; Godfrey,1932a : 275. (P)
Iambrix salsala salsala ; Evans,1949 : 272. (Siam)
Iambrix salsala salsala ; Shirôzu & Saigusa,1962 : 33-34, pl.2, figs.[11,♂,12,♂(Un), as Ancistroides nigrita ], Textfig.6,♂genitalia. (Doi Inthanon; Mae Hoi)
Iambrix salsala salsala ; Kawazoé,1964 : 126-127, pl.1, figs.7,♀,8,♀(Un). (N: Doi Suthep; Ping Kong / P: Khao Chong)
Iambrix salsala salsala ; Pinratana,1985 : 59, pl.20, figs.117,♂,♂(Un). (Chiang Mai; Chaiyaphum; Nakhon Nayok; Bangkok; Chanthaburi; Trang; Yala)
Iambrix salsala salsala ; Yamaguchi & Aoki,2000 : 250, fig.16,♂. (Mukdahan: Phu Mu; Ban Naudom; Phu Shang; Ubon Ratchathani: Kaeng Tana)
Iambrix salsala salsala ; Yamaguchi & Aoki,2002 : 12. (Ubon Ratchathani: Ban Na Pho Krang; Phu Chong Na Yoi; Kaeng Tana; Kaeng E-Keow; Phu Hin Dang; Mukdahan: Ban Lao Kram; Nam Tok Kham Cha I; Ban Na Lak; Ban Gok Tum)
Iambrix salsala salsala ; Yamaguchi, Aoki & Akiyama,2005 : 44. (Ubon Ratchathani: Kaen Tan; Nam Tok Soi Sawan; Phu Chong Na Yoi; Mukdahan: Ban Lao Kram; Ban Na Lak; Phu Hin Khan; Nam Tok Tad Ton)
Iambrix salsala salsala ; Katsuyama et al.,2005 : 63, fig.16,♂. (Umphang: Tha Sai; Umphang)
Iambrix salsala salsala ; RDPB & DNP,2006-2008 : Butterflies in Hala-Bala Forest, fig.[♀](Un). (Narathiwat: Hala Bala) (last update: 2007.01.02 ; accessed: 2023.07.12)
Iambrix salsala salsala ; Ek-Amnuay,[2007] : 782, pl.359, figs.H142,♂,♂(Un),♂,♂(Un),♀,♀(Un). (Ban Hong; Khao Yai; Trat)
Iambrix stellifer ; Ek-Amnuay,[2007] : 782, pl.359, figs.H143,♂[♀],♂[♀](Un). (Yala)
Iambrix salsala salsala ; Les Day,2011-2022 : Samui Butterflies, figs.[♀],[♀](Un). (Koh Samui) (accessed: 2022.02.11)
Iambrix salsala salsala ; Kimura et al.,2011 : 61, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao; Doi Suthep; Maethro; Saraburi; Nakohn Nayok; Sarika; Wan Nam Khiewo; Kao Soi Dao; Kratchin WF; Kao Saverb; Bang Phra; Kao Chamaoh; Doi Musso; Sai York Noi; Khlong Tariew; Kuwai Yai; Kaeng Krachang; Huey Yahn WF; Muang Shone; Khlong Naka; Ka Chong; Satul; Thaleban; Than To WF.)
Iambrix salsala salsala ; Ek-Amnuay,2012 : 836, pl.385, figs.H138,♂,♂(Un),♂,♂(Un),♀,♀(Un). (Ban Hong; Khao Yai; Trat)
Iambrix stellifer ; Ek-Amnuay,2012 : 836, pl.385, figs.H139,♀,♀(Un). (Yala)
Iambrix salsala salsala ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Iambrix salsala salsala ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Sa kaeo; Saraburi; Chanthaburi; Trat) (accessed: 2023.08.03)
Iambrix salsala salsala ; Küppers & Janikorn,2017 : 90-91, figs.Abb.H-125-130. (Phang Nga)
Iambrix salsala salsala ; Chaianunporn & Chaianunporn,2019 : 85. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Iambrix salsala salsala ; Motono & Negishi,1989 : 81, pl.114, figs.A1,♂,A2,♂(Un),A3,♀,A4,♀(Un). (Luang Prabang; Vang Vieng; Tha Ngone; Nam Moh; Vientiane; Ban Houei Sai)
Iambrix salsala salsala ; Osada, Uémura & Uehara,1999 : 223, pl.139, fig.♂(Un). (Luang Phrabang)
Iambrix salsala salsala ; Nimura & Wakahara,2005 : 49. (Namdhua; Thabeng; Thakhek)
Iambrix salsala salsala ; Onodera,2022 : 116, pl.252, figs.♂,♂(Un),♀,♀(Un). (Vientiane: Vang Vieng; Khammouan: Nahin)

[ CAMBODIA ]
Iambrix salsala salsala ; Shirôzu & Saigusa,1962 : 33-34, pl.2, fig.10,♂, Textfig.6,♂genitalia. (Puok)
Iambrix salsala ; Onodera,2007 : 21,22,23,28, fig.111♂[♀]. (Rattanakiri: Ou Chum; Ta Veaeng; Veun Sai; Siem Reap: Phnum Kuhlen)
Iambrix salsala salsala ; Kosterin,2019 : 100. (Koh Kong; Preah Sihanouk)
Iambrix salsala salsala ; Kosterin,2019b : 34. (Mondulkiri)
Iambrix salsala salsala ; Kosterin,2020 : 333. (Siem Reap; Preah Vihear; Stueng Treng)
Iambrix salsala salsala ; Chartier & Kosterin,2022 : 125. (Kep; Koh Kong; Mondulkiri; Preah Sihanouk; Preah Vihear; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Iambrix salsala vasuba ; Metaye,1957 : 78. (N)
Iambrix salsala salsala ; Inoué & Kawazoé,1966 ; 96, fig.89,♀(Un). (Phan Tho; Vo Dat; Trang Bom; Ba Diem; Lai Thieu; Col de Blao; Banmethuot)
Iambrix salsala salsala ; Devyatkin & Monastyrskii,1999 : 167. (Tam Dao; Na Hang; Cao Bang; Cuc Phuong; Hoa Binh; Ba Na; Pu Mat; Vu Quang; Bach Ma; Buon Luoi)
Iambrix salsala salsala ; Ikeda et al.,2001b : 63, figs.3:3,♂,4,♂(Un),5:22(♂genitalia). (Cuc Phuong)
Iambrix salsala ; Vu,2003 : 59. (Phu Quoc)
Iambrix salsala salsala ; Luong et al.,2004 : 14, 88, figs.♂,♂(Un). (Cuc Phuong)
Iambrix salsala ; Monastyrskii et al.,2006 : 45. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Iambrix salsala salsala ; Miyazaki, Saito & Saito,2007b : 11, figs.H-72,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Madagui; Dong Nai: Thac Mai)
Iambrix salsala salsala ; Monastyrskii & Devyatkin,2015 : 78. (N / C / S)
Iambrix salsala salsala ; Saito et al.,2017 : 76. (S: Nui Cam; Nui Ket)

[ THAILAND ]
Mae Hong Son : 7♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 9♂ 3♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Lampang : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phrae : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 7♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Kanchanaburi : 2♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chonburi : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Trat : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchaburi : 3♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Ranong : 10♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Surat Thani (Koh Phangan dist.) : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nakhon Si Thammarat : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Satun : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Yala : 9♂ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Narathiwat : 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is very common in secondary and montane forests at low to moderate elevations (Alt. 20 - 1300m) and is often found at forest edges and grasslands. Both sexes visit flowers.
Incredibly, I have seen natural hybridisation between this species and Arnetta atkinsoni in Chiang Mai. This is the only case of mating I have ever seen between two different genera.

Moore, F.,1866 : On the lepidopterous insects of Bengal.
Proc. zool. Soc. Lond. 1865(3):755-822, 3 pls.

Fruhstorfer, H.,1910 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 24(3-5):58-104.

♂, Yala, Thailand.	♀ + ♂ (mating), Chiang Mai, Thailand.
♂, Lampang, Thailand.	♂, Chiang Mai, Thailand.
Natural mating between Iambrix salsala and Arnetta atkinsoni, Chiang Mai, Thailand.

1999.09.04 - 2024.06.30.