Polyura arja arja  (Felder & Felder, [1867])

Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♀ Up. Polyura arja arja ♀ Un.
♂, Nan, Thailand. ♀, Nan, Thailand.
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ Up. Polyura arja arja ♂ Un.
♂, Nan, Thailand. ♂, Chonburi, Thailand.
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ genitalia Polyura arja arja ♂ genitalia
♂, Phetchaburi, Thailand. ♂ genitalia, Nan, Thailand.

arja ?
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ Up. Polyura arja arja ♂ Un.
♂, Chiang Mai, Thailand. ♂, Vientiane, Laos.
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ genitalia Polyura arja arja ♂ genitalia
♂, Chonburi, Thailand. ♂ genitalia, Vientiane, Laos.

f.roeberi
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ Up. Polyura arja arja ♂ Un.
♂, Tak, Thailand. ♀, Kachin, Myanmar.
Polyura arja arja ♂ Up. Polyura arja arja ♂ Un. Polyura arja arja ♂ genitalia Polyura arja arja ♂ genitalia
♂, Kachin, Myanmar. ♂ genitalia, Tak, Thailand.

ON : Charaxes arja Felder & Felder, [1867]
OD : Reise Fregatte Novara, Lep. Rhop. (3):438.
TL : India septentrionalis: Assam. (NHML)
Distribution : Sikkim to Myanmar, Thailand, Laos, Cambodia,
Vietnam, Yunnan.
Polyura arja arja map

Synonyms

Charaxes arja roeberi Fruhstorfer, 1898 : Ent. Nachr. 24(4):59. TL. Khasia Hills.
Eulepis arja f.temp.vernus Rothschild, 1899 : Novit. zool. 6:244-245. TL. Sikkim. (NHML)


Type Material Information

arja :
Lectotype ♂, India, Silhet. designated by R.L.Smiles,1979. (NHML)
Paralectotype 1♂, India Assam. designated by R.L.Smiles,1979.(NHML)
vernus : Holotype ♂, 23.Mar.1888. Sikkim. (NHML)


Records taken from Literature

[ THAILAND ]
Eriboea arja ; Fruhstorfer,1914 : 720. (Petchaburi)
Eulepis arja ; Godfrey,1916 : 133. (Chonburi: Sriracha)
Eriboea arja ; Godfrey,1927 : 124. (N)
Eriboea arja ; Godfrey,1930 : 301. (Prae: Me Song forest; Chiang Rai; Muak Lek; Chonburi: Sriracha; Me Wong)
Polyura arja arja ; Pinratana,1979 : 98, figs.N167,♂,♂(Un). (Chonburi; Nan)
Polyura arja arja ; Smiles,1984 : 177. (Pak Jong; Mae Wong; [Me Tharn]; Mae Sariang; Hin Lap; [Muok-Lek]; [Pang Yao]; [Raheng])
Polyura arja arja ; Pinratana & Eliot,1996 : 98, pl.75, figs.188,♂,♀. (Chiang Mai; Chaiyaphum; Phisanulok; Nakhon Ratchasima; Nakhon Nayok; Chonburi; Chanthaburi; Saraburi; Kanchanaburi)
Polyura arja arja ; Yamaguchi & Aoki,2000 : 256, fig.116,♂. (Mukdahan: Phu Pha Thaep; Phu Mu)
Polyura arja arja ; Yamaguchi & Aoki,2002 : 31. (Mukdahan: Ban Lao Kram; Ban Na Lak; Ubon Ratchathani: Kaeng E-Keow)
Polyura arja arja ; Yamaguchi, Aoki & Akiyama,2005 : 52. (Ubon Ratchathani: Phu Pung; Mukdahan: Ban Lao Kram)
Polyura arja arja ; Katsuyama et al.,2005 : 71, fig.178,♂. (Umphang: Tha-Sai; Umphang)
Polyura arja arja ; Katsuyama et al.,2006 : 50, fig.124,♂. (Nan: Ban Nam Poon)
Polyura arja arja ; Ek-Amnuay,[2007] : 486, pl.217, figs.N201,♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao)
Polyura arja arja ; Ek-Amnuay,2012 : 358, pl.152, figs.N189,♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao)
Polyura arja arja ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs, Early Stages. (Chiang Mai; Chanthaburi) (accessed: 2021.07.01)
Polyura arja arja ; Kimura et al.,2016 : 133, figs.♂,♂(Un),♀,♀(Un). (Chiang Mai; Maethro; Thung Saleng Luang; Phu Khiewo; Nakohn Nayok; Saraburi; Wan Nam Khiewo; Kao Soi Dao; Pliewn WF; Kao Khiewo; Sai York Noi)

[ LAOS ]
Eriboea arja f.vernus ; Dubois & Vitalis,1924 : 46. (Ban-Samoui)
Eriboea arja f.roeberi ; Dubois & Vitalis,1924 : 46. (Nam-Tie)
Polyura arja arja ; Motono & Negishi,1989 : 62, pl.77, figs.5,♀,6,♀(Un). (Vang Vieng; Phatong; Nam Moh; Tha Ngone; Vientiane; Luang Prabang; Ban Houei Sai)
Polyura arja arja ; Osada, Uémura & Uehara,1999 : 208, pl.78, figs.♂,♂(f.roeberi),♀,♀,♂(Un). (Oudomxay; Vientiane; Nong Het)
Polyura arja arja ; Nimura & Wakahara,2005 : 43-44. (Namdhua; Thakhek)
Polyura arja arja ; Onodera,2009b : 39. (Bolikhamxai: Thabok)
Polyura arja arja ; Nakamura & Wakahara,2012 : 47.

[ CAMBODIA ]
Polyura arja arja ; Woodfield & Murton,2006 : fig,♂. (Botum Sakor NP.)

[ VIETNAM ]
Eriboea arja ; Fruhstorfer,1914 : 720. (Tonkin: Chiem Hoa)
Eriboea arja ; Dubois & Vitalis,1919 : 233. (Tonkin)
Eriboea arja f.arya [sic] ; Dubois & Vitalis,1924 : 46. (Tonkin: Chiem-Hoa)
Eriboea arja roeberi ; Metaye,1958b : 174. (Dong Nai: Trang Bom)
Polyura arja arja ; Smiles,1984 : 177. (Chiem Hoa; [Nam-Hou]; [Ko-Tich]; Van Bu; Cha Pa)
Polyura arja arja ; Ikeda et al.,2000 : 37, figs.8:7,♂,8,♂(Un), fig.9:47(♂,genitalia). (Cuc Phuong)
Polyura arja ; Luong et al.,2004 : 8. (Cuc Phuong)
Polyura arja arja ; Miyazaki, Saito & Saito,2006b : 34, fig.N-112,♂. (Lam Dong: Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Polyura arja arja ; Miyazaki, Saito & Saito,2007c : 38, fig.N-112,♀. (Lam Dong: Dambri)
Polyura arja arja ; Nihira,2009 : 99, fig.♂. (Phu Quoc)
Polyura arja arja ; Monastyrskii & Devyatkin,2015 : 42. (N / C / S)
Polyura arja arja ; Monastyrskii,2019 : 4(1):87-88, pl.13, figs.2a♂,2b♂(Un),2c♀. (N: Muong Nhe; Huu Lien ; Cuc Phuong / C: Pu Mat; Nui Chua; Yok Don; Chu Yang Sin; Cat Tien / S: Cat Tien; Binh Chau)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 10♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 2♂ 4♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chonburi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Dong Nai : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


< arja ? >

[ THAILAND ]
Chiang Mai : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chonburi : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

[ LAOS ]
Vientiane : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


< f.roeberi >

[ THAILAND ]
Tak : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

[ MYANMAR ]
Kachin (N.E.Putao) : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is common in montane forest at low to moderate elevations (Alt.300m - 900m).
I suspect Polyura arja has several cryptic species. Further taxonomic study is needed.


Literature

Felder, C. & Felder, R.,1865-1875 :
Rhopalocera. In Reise der Osterreichischen Fregatte "Novara" um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Zweiter Band:
Abtheilung. Wien : vi + 548 + [1] pp in 5 parts, 140 pls.

Fruhstorfer, H.,1898 : Ueber einige Charaxes-Arten.
Ent. Nachr. 24(4):53-60.

Rothschild, L.W. & Jordan, K.,1899 : A monograph of Charaxes and the allied Prionopterous genera. Part 2.
Novit. zool. 6:220-286, pl.7.

Smiles, R.L.,1982 : The taxonomy and phylogeny of the genus Polyura Billberg (Lepidoptera: Nymphalidae)
Bull. Brit. Mus. Ent. (Nat. Hist.) 44(3):115-237. London.

Lang, S.Y.,2009 : A Study on the systematics of Nymphalidae (excepting Satyrinae) from China (Lepidoptera: Papilionoidea)
Beijing, P.R.China.

Lang, S.Y.,2012 : The Nymphalidae of China (Lepidoptera, Rhopalocera) Part 1
Libytheinae, Danainae, Calinaginae, Morphinae, Heliconiinae, Nymphalinae, Charaxinae, Apaturinae, Cyrestinae, Biblidinae, Limenitinae
456 pp, 32 col. + 41 black&white pls. Tshikolovets Publications, Pardubice.

Fang, L. & Zhang, Y.,2018 : Review of the genus Polyura Billberg (Lepidoptera: Nymphalidae) from China with description of a new species
Entomotaxonomia 40(1):27-45.


2006.12.09 - 2021.07.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.