Enispe euthymius sychaeus Brooks, 1949

Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un. Enispe euthymius sychaeus ♀ Up. Enispe euthymius sychaeus ♀ Un.
♂ (dsf), Chiang Mai, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un. Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un.
♂ (wsf), Phong Saly, Laos. ♂ (wsf), Phong Saly, Laos.
Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un. Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un.
♂ (dsf), Bolikhamxai, Laos. ♂ (wsf), Phong Saly, Laos.
Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un. Enispe euthymius sychaeus ♂ Up. Enispe euthymius sychaeus ♂ Un.
♂ (ds), Mae Hong Son, Thailand. ♂ (wsf), Chiang Mai, Thailand.
Enispe euthymius sychaeus ♂ genitalia Enispe euthymius sychaeus ♂ genitalia Enispe euthymius sychaeus ♀ genitalia Enispe euthymius sychaeus ♀ genitalia
♂ genitalia, Lampang, Thailand. ♀ genitalia, Fang, Chiang Mai, Thailand.

ON : Enispe euthymius sychaeus Brooks, 1949
OD : Entomologist 82: 259.
TL : 'Chapa, Cochin China' [Sapa, Lao Cai, N.Vietnam]. (NHML)
Distribution : Myanmar, S.E.Tibet, Yunnan, Thailand, Laos, N. & C.Vietnam.
● ssp.sychaeus
? unclear (E. euthymius or E. duranius)
ssp.congi
ssp.'congi' ??
Enispe euthymius sychaeus map

Records taken from Literature

[ THAILAND ]
Enispe euthymius euthymius f. tesselata ; Godfrey,1930 : 263. (Prae: Me Song forest)
Enispe intermedia intermedia ; Pinratana,1983 : 60, 64, pl.48, fig.29,♂. (Chiang Mai; 'Chaiyaphum') (partim)
Enispe intermedia intermedia ; Ek-Amnuay,[2007] : 344, pl.147, figs.A32a♂,♂(Un),♀. (Doi Pha Hom Pok; Wiang Pa Pao)
Enispe euthymius euthymius ; Inayoshi,2011 - 2023 : Butterflies in Indo-China, figs.♂,♂(Un),♀,♀(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un). (Chiang Rai; Chiang Mai; Lampang) (30.Jun.2023 last update. 31.Aug.2023 changed.)
Enispe duranius intermedia ; Ek-Amnuay,2012 : 348, pl.147, figs.N179a♂,♂(Un),♀. (Doi Pha Hom Pok; Wian Pa Pao)
Enispe euthymius euthymius ; Kimura et al.,2016 : 154, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Doi Par Hom Pok; Doi Chiang Dao; Doi Suthep; Ban Salok)

[ LAOS ]
Enispe euthymius f. tesselata [sic] ; Dubois & Vitalis,1924 : 34. (Muong-Om; Luang-Prabang; Muong-You; Tam-Sa; Ban Thao-Phan; Pou-So-Ko; Ban-Na-Mone)
Enispe euthymius corbeti ; Motono & Negishi,1989 : 67, pl.93, figs.7,♂,8,♂(Un). (Nam Moh)
Enispe euthymius sychaeus ; Osada, Uémura & Uehara,1999 : 212, pl.102, fig.♂. (Lak Sao)
Enispe euthymius sychaeus ; Uémura,2005 : 5, fig.29,map.
Enispe euthymius sychaeus ; Nakamura & Wakahara,2012 : 50.
Enispe euthymius sychaeus ; Nakamura,2021 : 9,13, figs.23,♂,23v♂(Un),23a(♂,genitalia). (Bolikhamxai: Nahin)
Enispe euthymius sychaeus ; Onodera,2022 : 66, pl.196, figs.♂,♂(Un),♂,♂(Un). (Xiengkhouang: Tha; Thna)

[ VIETNAM ]
Enispe euthymius f. tesselata [sic] ; Dubois & Vitalis,1924 : 34. (Tonkin) (partim)
Enispe duranius ; Lemée,1950 : 9. (Hagiang)
Enispe euthymius tesselatus ; Metaye,1957 : 100. (N / C)
Enispe euthymius ; Monastyrskii et al.,2006 : 40. (Thua Thien Hue: Bach Ma)
Enispe euthymius sychaeus ; Monastyrskii,2011 : 122-123, pl.30, fig.2(HT,♂). (N: Hoang Lien; Van Ban; Ba Be / C: Xuang Lien; Pu Hoat; Pu Mat; Huong Son)
Enispe euthymius sychaeus ; Monastyrskii & Devyatkin,2015 : 30. (N / C: to 'Gia Lai')
Enispe euthymius sychaeus ; Nakamura,2021 : 13, figs.24,♂,24v♂(Un),24a(♂,genitalia). (N: Ha Giang)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 1♂ (genitalia examined).
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Rai : 1♂ (genitalia examined).
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai (excluding Omkoi dist.) : 7♂ 1♀ (7♂ genitalia examined, 1♀ genitalia dissected).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 5♂ (1♂ genitalia dissected, 4♂ examined).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Luang Phabang : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Phong Saly : 3♂ (genitalia examined).
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 1♂ (genitalia examined).
JanFebMarAprMayJun JulAugSepOctNovDec

Oudomxai : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Xieng Khouang : 2♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The distribution map of duranius and euthymius
Enispe duranius-euthymius distribution


The butterfly is not uncommon in montane forests at moderate to high elevations (Alt. 1,000–1,800 m).
According to Brooks (1949: 258) and Nakamura (2021: 9), it occurs sympatrically with Enispe duranius in parts of Myanmar and Laos. Previously, the distribution of this species within Thailand was considered limited to the area around Chiang Mai city and eastwards; however, in 2026, its presence was confirmed in Mae Hong Son Province. The distributional ranges of these two species are therefore thought to overlap between Mae Hong Son and the area west of Chiang Mai city. Although no instances of strict sympatry at the exact same locality have yet been confirmed, it is expected to be recorded in the future.
This species is distinguishable from E. duranius solely by examination of the genitalia. Consequently, records unsupported by genitalic dissection cannot be reliably utilized. Although several records exist from Chaiyaphum and Phetchabun provinces in north-central Thailand, the lack of genitalic examination leaves their specific identity uncertain; they could represent either this species or E. duranius. Therefore, further taxonomic and distributional studies are needed.
The populations from Yunnan to N. Thailand are morphologically much closer to ssp. sychaeus than to the nominotypical subspecies, as Huang (2023: 103-104) pointed out.

Differences in the genitalia of euthymius and duranius
Enispe euthymius sychaeus ♂ genitalia Enispe duranius intermedia ♂ genitalia Enispe euthymius sychaeus ♀ genitalia Enispe duranius intermedia ♀ genitalia
♂, Enispe euthymius
(Lampang)
♂, Enispe duranius
(Tak)
♀, Enispe euthymius
(Chiang Mai: Fang)
♀, Enispe duranius
(Chiang Mai: Chom Thong)


Literature

Brooks, C.J.,1949 : New subspecies in the Genera Faunis, Aemona, Stichophthalma, and Enispe, with Revisional Notes (Lep. Amathusiidae).
Entomologist 85: 256-259.

Nakamura, N.,2021 : Notes on the butterflies of Laos (XII):
New findings on Amathusiini (Morphinae: Nymphalidae) including description of a new subspecies of Faunis excelsa (Fruhstorfer, 1901).
Butterflies 85: 4-14.

Huang, H.,2023 : New or little known butterflies from China - 5 (Lepidoptera: Papilionidae, Pieridae, Nymphalidae, Lycaenidae et Hesperiidae).
Neue ent. Nachr. 82: 93-214.


Enispe euthymius sychaeus ♂ Enispe euthymius sychaeus ♂
♂, Chiang Mai, Thailand. ♂, Chiang Mai, Thailand.
Enispe euthymius sychaeus ♂ Enispe euthymius sychaeus ♂
♂, Lampang, Thailand.
Enispe euthymius sychaeus ♂ Enispe euthymius sychaeus ♂
♂, Mae Hong Son, Thailand.

2007.10.10 - 2026.06.30.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.