Ypthima savara  Grose-Smith,1887

Ypthima savara ♂ Up. Ypthima savara ♂ Un. Ypthima savara ♀ Up. Ypthima savara ♀ Un.
♂ (dsf), Phrae, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Ypthima savara ♂ Up. Ypthima savara ♂ Un. Ypthima savara ♀ Up. Ypthima savara ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Ypthima savara ♂ Up. Ypthima savara ♂ Un. Ypthima savara ♂ Up. Ypthima savara ♂ Un.
♂, Phong Saly, Laos. ♂, Luang Pabang, Laos. (photo by H.Onodera)
Ypthima savara ♂ Up. Ypthima savara ♂ Un. Ypthima savara ♀ Up. Ypthima savara ♀ Un.
♂, Yala, Thailand. ♀, Yala, Thailand.
Ypthima savara ♂ Up. Ypthima savara ♂ Un. Ypthima savara ♀ Up. Ypthima savara ♀ Un.
♂, Phetchabun, Thailand. ♀ (dsf), Chiang Mai, Thailand.

ON : Yphthima [sic] savara Grose-Smith,1887
OD : Ann. Mag. nat. Hist. (5)20:267.
TL : Burmah, Siamese frontier. (NHML)
Distribution : Myanmar, Thailand, Laos, Vietnam, W.Malaysia.

Synonym

Ypthima tonkiniana Fruhstorfer,1908 : Ent. Zeit. 22(31):127. TL. Chiem Hoa, N.Vietnam. (NHML)


Type Materials Information

savara ; Syntype 1♂, Rh3410, 1♂, Rh3411, Burmah, Siamese frontier. (NHML)
tonkiniana ; Syntype 1♀, #141995, Chiem Hoa, Tonkin. Aug.Sept. (NHML)


Records taken from Literature

[ THAILAND ]
Ypthima savara savara ; Godfrey,1927 : 121. (N)
Ypthima savara savara ; Godfrey,1930 : 244. (Chiang Rai: Doi Chang / Khun Than / Prae: Me Song forest / Umphang)
Ypthima savara savara ; Aoki & Uémura,1984 : 83. figs.11,a-b. (Mae Hong Son: Mae Hong Son / Chiang Mai: Tha Tong ; Tam Tab Tao ; Chiang Dao ; Mae Sa ; Muang Ngai ; Nam Tok Mae Klang / Chiang Rai: Wiang Pa Pao / Tak: Doi Musso / Kanchanaburi: Sai Yok Noi / Phitsanulok: Tong Salang Luang / Loei: Phu Kadung ; Nam Nao)
Ypthima savara savara ; Pinratana,1988 : 33, pl.34, figs.92,♂,♀(Un). (Chiang Mai / Loei / Phitsanulok / Nakhon Ratchasima / Kanchanaburi / Tak)
Ypthima savara savara ; Yamaguchi & Aoki,2000 : 254, fig.79,♂(Un). (Mukdahan: Phu Pha Thaep)
Ypthima savara savara ; Yamaguchi & Aoki,2002 : 24. (Mukdahan: Ban Lao Kram ; Ban Na Lak ; Phu Huay Sing ; Ban Gok Tum / Ubon Ratchathani: Ban Na Pho Krang ; Phu Chong Na Yoi)
Ypthima savara savara ; Yamaguchi, Aoki & Akiyama,2005 : 49. (Mukdahan: Ban Lao Kram ; Ban Na Lak / Ubon Ratchathani: Nam Tok Soi Sawan)
Ypthima savara savara ; Katsuyama et al.,2005 : 68, fig.128,♂. (Umphang: Tha-Sai)
Ypthima savara savara ; Katsuyama et al.,2006 : 49, fig.98,♂. (Nan: Ban Nam Poon)
Ypthima fasciata torone ; Ek-Amnuay,[2007] : pl.121, figs.S94,♂(top-left),♂(top-centerUn),♀(bottom),♀(bottom,Un). (in part)
Ypthima savara savara ; Ek-Amnuay,[2007] : 298, pl.121, figs.S96,♂,♂(Un),♀[♂],♀[♂](Un). (Chiang Mai / Phu Khieo)
Ypthima savara savara ; Ek-Amnuay,2012 : 298, pl.121, figs.N131,♂,♂(Un),♂(Un),♀,♀(Un). (Chiang Mai / Phu Khieo / Kaeng Krachan)
Ypthima savara savara ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Ypthima savara savara ; Kimura et al.,2016 : 194, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao / Doi Suthep / Thung Saleng Luang / Phu Kradung / Phu Khiewo / Wan Nam Khiewo / Ban Na Lak / Kratching WF / Kaeng Chai / Kao Phang WF / Sai York Noi / Kuwai Yai / Khlong Naka)
Ypthima savara savara ; Saito & Inayoshi,2018 : 30, figs.21,♂,♂(Un),22,♀,♀(Un). (Yala)
Ypthima savara savara ; Chaianunporn & Chaianunporn,2019 : 87. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Ypthima savara savara ; Motono & Negishi,1989 : 66, pl.91, figs.7,♂,8,♂(Un). (Vang Vieng)
Ypthima sarava savara ; Osada, Uémura & Uehara,1999 : 211, pl.96, figs.♂,♀(Un). (Thabok / Vientiane)
Ypthima sarava tonkiniana ; Uémura,2005 : 1, figs.4(map), 33,♂(wsf),34,♂(dsf). (Boun Neua / Thabok / Vieng Thong / Ban Boung Kuang / Lak Sao)
Ypthima sarava tonkiniana ; Uémura,2006 : 7, fig.39,map. (Vieng Thong)
Ypthima sarava tonkiniana ; Uémura,2007 : 3, fig.21,map. (Nakhai to Lak Sao)
Ypthima savara savara ; Onodera,2009b : 41. (Khammouan: Nacay)
Ypthima sarava tonkiniana ; Uémura,2011 : 3. (C: Phon Si ; Nahin Pass ; Nakhai to Lak Sao 20km ; near Nam Teun)
Ypthima sarava tonkiniana ; Nakamura & Wakahara,2012 : 49.

[ CAMBODIA ]
Ypthima savara f.savara ; Dubois & Vitalis de Salvaza,1924 : 27. (Pnom-Penh)
Ypthima savara savara ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)

[ VIETNAM ]
Ipthima [sic] tonkiniana ; Dubois & Vitalis de Salvaza,1919 : 222. (Tonkin)
Ypthima savara f.tonkiniana ; Dubois & Vitalis de Salvaza,1924 : 27. (Tonkin: Chiem-Hoa)
Ypthima savara tonkiniana ; Metaye,1957 : 102. (N)
Ypthima savara savara ; Uémura & Monastyrskii,2004 : 24-25, figs.2i, 5.1,♂. (C: Phong Dien ; Bach Ma ; Ba Na ; Kon Ka Kinh ; Ban Don ; Bao Loc / S: Nghia Trung ; Cat Tien)
Ypthima savara tonkiniana ; Uémura & Monastyrskii,2004 : 24, figs.2g-h, 5.2,♂. (N: Ba Be / C: Ben En ; Pu Mat)
Ypthima savara savara ; Monastyrskii,2005 : 110-111. (C: Phong Dien ; Bach Ma ; Kon Ka Kinh ; Ban Don ; Cat Tien ; Da Teh / S: La Nga ; Nghia Trung)
Ypthima savara tonkiniana ; Monastyrskii,2005 : 110-111, pl.29, figs.1a,♂,1b,♂(Un),1c,♀,1d,♀(Un). (N: Na Hang ; Ba Be / C: Ben En ; Pu Mat)
Ypthima savara savara ; Miyazaki, Saito & Saito,2006c : 34, fig.S-33,♀[♂]. (Lam Dong: Dambri)
Ypthima savara savara ; Miyazaki, Saito & Saito,2007c : 39, fig.S-33,♀. (Lam Dong: Dambri)
Ypthima savara savara ; Monastyrskii & Devyatkin,2015 : 26. (C: to Lam Dong)
Ypthima savara tonkiniana ; Monastyrskii & Devyatkin,2015 : 26. (N / C: to Quang Tri)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 11♂ 11♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 1♂ 1♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Yala : 3♂ 1♀ (incl.photo by S.Atdhabhan).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Luang Phabang : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Phong Saly : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Oudomxai : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 2♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Vientiane : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Sekong : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
Literature

Grose-Smith, H.,1887 : Descriptions of eight new Species of Asiatic Butterflies.
Ann. Mag. nat. Hist. (5)20:265-268.

Fruhstorfer, H.,1908 : Neue ostasiatische Satyriden
Ent. Zeit. 22(31):127.

Uémura, Y.,2020 : A synonymic list of the sakra-group of the genus Ypthima Hubner (Lepidoptera: Nymphalidae, Satyrinae)
Bull. Toyosato Mus. Ent. (30):1-20.


Ypthima savara ♂ Ypthima savara ♀
♂ (dsf), Chiang Mai, Thailand. ♀ (dsf), Phetchabun, Thailand.
Ypthima savara ♂ Ypthima savara ♂+♀
♂, Chiang Mai, Thailand. ♂+♀, Chiang Mai, Thailand.
Ypthima savara ♂ Ypthima savara ♀
♂ (dsf), Chiang Mai, Thailand. ♂ (dsf), Yala, Thailand. (photo by S.Atdhabhan)

1998.12.08 - 2020.12.31.
A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.