♂ (dsf), Phrae, Thailand.		♀ (dsf), Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.

ON : Calysisme intermedia Moore, [1892] OD : Lep. Ind. 1:187-188, pl.60, fig.2. TL : Tenasserim. (NHML) Distribution : Assam to Myanmar, Thailand, Laos, Cambodia, Vietnam.

Mycalesis visala neovisala Fruhstorfer, 1908 : Verh. zool.-bot. Ges. Wien. 58(6/7):158. TL. Than Moi ; Chiem-Hoa, Tonkin.
Mycalesis perseoides kashia Evans, 1912 : J. Bombay nat. Hist. Soc. 21(2):558, 568. TL. Khasi Hills.

intermedia:
Syntype 1♂ (wsf), Rh2963, Tenasserim. (NHML) ; Syntype 1♂ (dsf), Rh2697, Tenasserim. (NHML) ;
Syntype 1♀ (dsf), Rh2968, Tenasserim. (NHML)
neovisala: Syntypes 3♂ (wsf), Than-Moi, Tonkin ; Syntypes 2♂ 1♀(dsf), Chiem Hoa, Tonkin.

[ THAILAND ]
Mycalesis intermedia intermedia ; Aoki & Yamaguchi,1984 : 50-51, figs.7-8(♂ genitalia),13(map),25,♂. (Mae Hong Son: Mae Hong Son; Chiang Mai: Fang; Tam Tab Tao; Doi Suthep; Chiang Rai: Wing Pa Pao; Phitsanulok; Tong Slang Luang; Loei: Phu Kadeng; Ban Pha Nok Khao; Khon Kaen: Nam Nao; Tak: Doi Musso; Nakhon Nayok: Khao Yai; Chanthaburi: Pong Nam Ron; Kanchanaburi: Sai Yok Noi) (in part)
Mycalesis intermedia ; Pinratana,1988 : 22, pl.24, figs.53,♂,♀(Un). (Chiang Mai; Ratchaburi; Chaiyaphum; Chanthaburi) (in part)
Mycalesis distanti mucianus ; Pinratana,1988 : 22. (Chiyaphum; Ratchaburi; Nakhon Nayok) (in part)
Mycalesis intermedia ; Yamaguchi & Aoki,2000 : 253, fig.74,♂(Un). (Mukdahan: Phu Sa Duak Bua; Phu Mu; Phu Pha Thaep; Ban Naudom; Phu Shang)
Mycalesis intermedia ; Yamaguchi & Aoki,2002 : 23. (Mukdahan: Ban Lao Kram; Nam Tok Tad Ton; Nam Tok Kham Cha I; Ubon Ratchathani: Phu Chong Na Yoi; Phu Hin Dang)
Mycalesis intermedia ; Yamaguchi, Aoki & Akiyama,2005 : 48. (Mukdahan: Ban Lao Kram; Nam Tok Tad Ton; Ubon Ratchathani: Nam Tok Soi Sawan; Phu Chong Na Yoi; Phu Pung)
Mycalesis intermedia ; Katsuyama et al.,2006 : 49, fig.94,♂. (Nan: Ban Nam Poon)
Mycalesis perseus cepheus ; Ek-Amnuay,[2007] : pl.106, figs.S57b♀,♀(Un). (in part)
Mycalesis mineus mineus ; Ek-Amnuay,[2007] : 268, pl.106, figs.S58a♂,♂(Un). (Khao Yai; Khao Soi Dao)
Mycalesis thailandica ; Ek-Amnuay,[2007] : 270, pl.107, figs.S60,♂,♂(Un),♀,♀(Un). (Kanchanaburi)
Mycalesis perseus cepheus ; Ek-Amnuay,2012 : pl.105, figs.N92b♂('Trang'),♂(Un,'Trang'),♀,♀(Un). (in part)
Mycalesis mineus mineus ; Ek-Amnuay,2012 : pl.106, figs.N93a♂,♂(Un,). (in part)
Mycalesis intermedia ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Mycalesis intermedia ; Kimura et al.,2016 : 180, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Par Hom Pok; Chiang Dao; Doi Suthep; Chiang Mai; Chan Phui; Doi Pa Chang; Thung Saleng Luang; Phu Khiewo; Saraburi; Nakohn Nayok; Wan Nam Khiewo; Kao Khiewo; Pliewn WF; Kao Saverb; Kao Soi Dao; Doi Musso; Thug Yai; Sai York Noi; Kanchanaburi; Khlong Tariwe; Kaeng Krachang)
Mycalesis intermedia ; Chaianunporn & Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)
Mycalesis intermedia ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2019- : Butterflies of Thailand, figs. (Saraburi; Nakhon Nayok; Chachoengsao; Ratchaburi) (accessed: 2023.07.01)

[ LAOS ]
Mycalesis intermedia intermedia ; Motono & Negishi,1989 : 65, pl.87, figs.1,♂,2,♂(Un),3,♀,4,♀(Un). (Luang Prabang; Nam Po; Nam Moh; Tha Ngone; Vientiane; Phatong)
Mycalesis intermedia ; Osada, Uémura & Uehara,1999 : 211, pl.94, fig.♂. (Vang Vieng)
Mycalesis intermedia ; Uémura,2004 : 3-4, fig.20,map, fig.34,♂(wsf), 36,♂(dsf), 38,♂(dsf). (Nam Tha; Oudomxay; Vang Vieng; Tha Ngone; Xiang Khouang; Ban Boung Kuang; Nha Hia; Lak Sao; Thakhek)
Mycalesis intermedia ; Nimura & Wakahara,2005 : 44. (Namdhua; Thakhek)
Mycalesis intermedia ; Uémura,2006 : 6, fig.29,map. (Phou Phanang; Tha Ngone; Muang Kham; Phong Si; Vieng Thong; Nha Hin; Nha Hin Pass; Lak Sao; Nam Kata; Nak Hai)
Mycalesis intermedia ; Uémura,2007 : 3, fig.17,map. (Phon Si; Nahin Pass; Mamkata; Nakhai to Lak Sao; Ban Lat Khoay)
Mycalesis intermedia ; Onodera,2009b : 41,43. (Khammouan: Nacay; Bolikhamxai: Phon Si)
Mycalesis intermedia ; Uémura,2011 : 2-3. (C: Tha Ngone; Ban Nakha; Phon Si; near Nahin; Nahin Pass; Nakhai to Lak Sao 20km; near Lak Sao; Namkata; near Phin; Phin-Tat Hai; near border to Vietnam)
Mycalesis intermedia ; Nakamura & Wakahara,2012 : 49, pl.8, figs.Sa-62,♀,♀(Un). (Vang Vieng)
Mycalesis intermedia ; Onodera,2022 : 58, pl.175, figs.♂(wsf),♂(wsf,Un),♂(wsf),♂(wsf,Un),♂(wsf),♂(wsf,Un),♀(wsf),♀(wsf,Un),♀(wsf),♀(wsf,Un),♀(wsf),♀(wsf,Un), pl.176, figs.♂(dsf),♂(dsf,Un),♂(dsf),♂(dsf,Un),♂(dsf),♂(dsf,Un),♀(dsf),♀(dsf,Un),♀(dsf),♀(dsf,Un),♀(dsf),♀(dsf,Un). (Khammouan: Nakai; Thakhek; Natung; Savannakhet; Bolihamxai: Phon Si; Xiengkouang: Tha; Luang Phabang; Phongsali: Sin Xai; Muang Mai)

[ CAMBODIA ]
Mycalesis intermedius ; Onodera,2007 : 22, fig.40,♂. (Rattanakiri: Ta Veaeng)
Mycalesis intermedia ; Onodera,2008 : 61. (Mondolkiri: Bousra)
Mycalesis intermedia ; Chartier & Kosterin,2022 : 116. (Kep; Koh Kong; Mondulkiri; Ratanakiri; Siem Reap)

[ VIETNAM ]
Mycalesis visala neovisala ; Fruhstorfer,1911 : 346. (Tonkin: Than Moi; Chiem Hoa)
Mycalesis intermedia ; Vitalis,1919 : 223. (Tonkin)
Mycalesis neovisala ; Vitalis,1919 : 223. (Tonkin)
Mycalesis visala f. visala ; Dubois & Vitalis,1924 : 30. (Tonkin: Chiem-Hoa; Than-Moi)
Mycalesis visala neovisala ; Lemée,1950 : 8. (Hagiang; Backan)
Mycalesis visala visala ; Metaye,1957 : 103. (N)
Mycalesis visala neovisala ; Metaye,1957 : 103. (N)
Mycalesis intermedia ; Luong et al.,2004 : 4, 49, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Mycalesis intermedia ; Monastyrskii,2005 : 91-92, pl.25, figs.1a♂(dsf),1b♂(dsf,Un),1c♂(wsf),1d♂(wsf,Un),1e♀. (N: Muong Nhe; Na Hang; Ba Be; Cat Ba; Cuc Phuong; Pu Luong / C: Pu Hoat; Pu Mat)
Mycalesis intermedia ; Miyazaki, Saito & Saito,2006c : 31, figs.S-29,♂,♀. (Lam Dong: Dambri; Gung Re; Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Mycalesis intermedia ; Nihira,2009 : 100. (Phu Quoc)
Mycalesis intermedia ; Monastyrskii & Devyatkin,2015 : 25. (N / C / S)
Mycalesis intermedia ; Saito et al.,2017 : 82. (S: Nui Cam)

[ THAILAND ]
Mae Hong Son : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 12♂ 8♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phrae : 4♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 1♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Kanchanaburi : 1♂ 3♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Luang Phabang : 1♂ 1♀ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Xieng Khouang : 1♀ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Bolikhamxai : 1♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is very common in secondary and montane forests at low to moderate elevations (Alt. 100 - 1000m).
Identification of this species requires examination of the colour and shape of the male androconial brand on the UnF and UpH, or the male genitalia. Most species in the mineus group are very difficult to identify using the underside markings, and the dry season forms are almost impossible to identify.

The differences of the Mycalesis mineus group. (♂ androconial brand). Both characteristics must be confirmed for identification.
	perseus	perseoides	mineus	intermedia	distanti	visala
♂ UnF
	black (small)	dark brown	dark brown (small)	dark brown (long)	brown	pale yellow (long)
♂ UpH
	black (small)	black (long and slender)	salmon pink or dark brown	pale yellow (oval, large)	pale yellow (large) (extends to the base of space 6)	pale yellow (long and slender)

Moore, F.,1890-1892 : Lepidoptera Indica. vol.I. Rhopalocera, Family Nymphalidae.
1:1-192, 1890 ; 193-176, 1891 ; 177-310, 1892. L.Reeve & Co., London.

Fruhstorfer, H.,1908 : Neue indo-australische Mycalesis und Besprechung verwandter Formen.
Verh. zool.-bot. Ges. Wien. 58(2/3):126-128; (6/7):129-239, pl.1.

Evans, W.H.,1912 : A list of Indian butterflies.
J. Bombay nat. Hist. Soc. 21(2):553-584, 21(3):969-1008.

Uémura, Y.,1985 : Identification guide to the Thai-Malayan species of the mineus-group of Mycalesis (Lepidoptera: Satyridae)
Yadoriga 123:2-9. (in Japanese)

♂ (dsf), Phrae, Thailand.	♂, Chiang Mai, Thailand.

1998.12.05 - 2023.07.31.