Lethe europa niladana  Fruhstorfer,1911

Lethe europa niladana ♂ Up. Lethe europa niladana ♂ Un. Lethe europa niladana ♀ Up. Lethe europa niladana ♀ Un.
♂, Chiang Mai, Thailand. ♀, Kanchanaburi, Thailand.
Lethe europa niladana ♂ Up. Lethe europa niladana ♂ Un. Lethe europa niladana ♀ Up. Lethe europa niladana ♀ Un.
♂, Chiang Mai, Thailand. ♀, Chanthaburi, Thailand.

ON : Lethe europa niladana Fruhstorfer,1911
OD : in Seitz, Grossschmett. Erde 9:315.
TL : Kumaon to Myanmar.
Distribution : India to Myanmar, Thailand, Laos, Cambodia, Vietnam, Yunnan.

Synonym

Lethe europa niladana f.gada Fruhstorfer,1911 : in Seitz, Grossschmett. Erde 9:316. TL. Muok Lek, Siam. (NHML)


Records taken from Literature

[ THAILAND ]
Lethe europa niladana f.gada ; Fruhstorfer,1911 : 316. (Siam)
Lethe europa niladana ; Godfrey,1916 : 121.
Lethe europa niladana ; Godfrey,1927 : 121. (N)
Lethe europa niladana ; Godfrey,1930 : 245.
Lethe europa malaya ; Reeves,1966 : 14. (Nakhon Nayok: Khao Yai)
Lethe europa niladana ; Pinratana,1988 : 12, pl.10, fig.18a,♂,♀,♂(Un). (Chiang Mai / Chanthaburi / Chonburi / Kanchanaburi / Bangkok / Ratchaburi)
Lethe europa niladana ; Yamaguchi & Aoki,2000 : 253, fig.71♂. (Mukdahan: Phu Mu ; Phu Pha Thaep)
Lethe europa niladana ; Yamaguchi & Aoki,2002 : 21. (Mukdahan: Phu Huay Sing ; Nam Tok Tad Ton ; Nam Tok Kham Cha I ; Ban Lao Kram ; Ban Na Lak / Ubon Ratchathani: Nam Tok Soi Sawan)
Lethe europa niladana ; Yamaguchi, Aoki & Akiyama,2005 : 48. (Mukdahan: Ban Lao Kram ; Phu Hin Khan / Ubon Ratchathani: Phu Pung)
Lethe europa niladana ; Katsuyama et al.,2005 : 68, fig.114♂. (Umphang: Tha-Sai ; Umphang)
Lethe europa niladana ; Katsuyama et al.,2006 : 48, fig.85♂. (Nan: Ban Nam Poon)
Lethe europa niladana ; Ek-Amnuay,[2007] : 240, pl.92, fig.S19a,♂,♂(Un),♀,♀(Un). (Tham Than Rod)
Lethe europa niladana ; Ek-Amnuay,2012 : 240, pl.92, fig.N55a,♂,♂(Un),♀,♀(Un). (Tham Than Lod)
Lethe europa niladana ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Lethe europa niladana ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, fig. (Ratchaburi / Chaiyaphum / Phetchabun)
Lethe europa niladana ; Kimura et al.,2016 : 167-168, fig.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Suthep / Ban Salok / Phu Kradung / Phu Khiewo / Sarika / Saraburi / Bang Sarrey / Doi Musso / Sai York Noi / Erawan WF / Kaeng Krachang / Ratchaburi)

[ LAOS ]
Lethe europa f.niladana ; Dubois & Vitalis de Salvaza,1924 : 28. (Laos)
Lethe europa f.beroe ; Dubois & Vitalis de Salvaza,1924 : 28. (Giranville)
Lethe europa niladana ; Motono & Negishi,1989 : 64, pl.82, figs.5♂,6♂(Un),7♀,8♀(Un). (Vang Vieng / Vientiane / Nam Moh / Luang Prabang)
Lethe europa niladana ; Osada, Uémura & Uehara,1999 : 210, pl.90, figs.♂,♂(Un). (Thabok / Oudomxay)
Lethe europa niladana ; Uémura,2003 : 3, fig.19,map. (Oudomxay / Vang Vieng / Ban Keun / Tha Ngone / Thabok / Lak Sao)
Lethe europa niladana ; Uémura,2006 : 2, fig.8,map. (Tha Ngone / Vieng Thong)
Lethe europa niladana ; Uémura,2007 : 1, fig.3,map. (Tad Thevada)
Lethe europa niladana ; Uémura,2011 : 1-2. (C: Tad Moon)
Lethe europa niladana ; Nakamura & Wakahara,2012 : 48.

[ CAMBODIA ]
Lethe gada ; Dubois & Vitalis de Salvaza,1919 : 223. (Cambodge)
Lethe europa niladana ; Woodfield & Murton,2006 : figs,♀,♀(Un). (Botum Sakor NP.)
Lethe europa gada ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)

[ VIETNAM ]
Lethe europa ; Joannis,1901 : 317. (Tonkin: Cao Bang)
Lethe europa niladana f.gada ; Fruhstorfer,1911 : 316. (Tonkin)
Lethe europa niladana ; Godfrey,1919 : 466. (S: Daban)
Lethe gada ; Dubois & Vitalis de Salvaza,1919 : 223. (Tonkin)
Lethe europa f.gada ; Dubois & Vitalis de Salvaza,1924 : 28. (Tonkin)
Lethe europa ; Lemee,1950 : 7-8. (Hagiang / Hanoi)
Lethe europa niladana f.gada ; Metaye,1957 : 102. (N)
Lethe europa gada ; Hao et al.,2004 : 4, 46, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Lethe europa gada ; Monastyrskii,2005 : 51-52, pl.12, figs.1a♂,1b♂(Un). (N: Tam Dao ; Ba Be ; Cuc phuong ; Cao Bang ; Lao Kai / C: Ben En ; Pu Mat ; Xuan Lien / S: Lo Go Sa Mat ; Cat Tien ; Saigon ; La Nga)
Lethe europa niladana ; Miyazaki, Saito & Saito,2006c : 28, figs.S-11♂. (Lam Dong: Dambri / Dong Nai: Nam Cat Tien / Bin Thuan: Dong Tien)
Lethe europa niladana ; Miyazaki, Saito & Saito,2007c : 39, fig.S-11♀. (Lam Dong: Dambri)
Lethe europa niladana ; Monastyrskii & Devyatkin,2015 : 22. (N / C / S)
Lethe europa niladana ; Saito et al.,2017 ; 81-82. (S: Nui Cam / Nui Coto)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 2♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 6♂ 1♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lamphun : 1♀ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 2♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 2♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Kanchanaburi : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 2♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Luang Prabang : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
Lethe europa niladana ♂ Lethe europa niladana ♀
♂, Chiang Mai, Thailand. ♀, Lamphun, Thailand.
Lethe europa niladana ♂ Lethe europa niladana ♂
♂, Lampang, Thailand. ♂, Phrae, Thailand.

1998.12.10 - 2018.10.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © Y.Inayoshi.