Elymnias malelas (Hewitson, 1863)

Elymnias malelas ♂ Up. Elymnias malelas ♂ Un. Elymnias malelas ♀ Up. Elymnias malelas ♀ Un.
♂, Mae Hong Son, Thailand. ♀, Nan, Thailand.
Elymnias malelas ♂ Up. Elymnias malelas ♂ Un. Elymnias malelas ♀ Up. Elymnias malelas ♀ Un.
♂, Nan, Thailand. ♀, Chanthaburi, Thailand.
Elymnias malelas ♂ Up. Elymnias malelas ♂ Un. Elymnias malelas ♀ Up. Elymnias malelas ♀ Un.
♂, Khammouan, Laos. (photo by K.Saito) ♀, Vinh Phuc, Vietnam. (photo by K.Saito)
Elymnias malelas ♂ Up. Elymnias malelas ♂ Un. Elymnias malelas ♀ Up. Elymnias malelas ♀ Un.
♂, Lam Dong, Thailand. (photo by K.Saito) ♀, Mae Hong Son, Thailand.
Elymnias malelas ♂ Up. Elymnias malelas ♂ Un. Elymnias malelas ♂ Up. Elymnias malelas ♂ Un.
♂, 2010.08.19. Surat Thani, Thailand. (photo by K.Saito) ♂, 2017.03.21. Yala, Thailand. (photo E.Ikari)

ON : Melanitis malelas Hewitson, 1863
OD : Ill. exot. Butts. 4:[70], pl.[36], figs.6,7,♂.
TL : 'East India' ["Sylhet" [southern Khasia Hills, India ;
north of Sylhet, Bangladesh]]. (NHML)
Distribution : Sikkim to Myanmar, Thailand, Laos,
Cambodia, Vietnam, Langkawi, W.Malaysia.
Elymnias malelas map

Synomyms

Elymnias malelas fa. subdecorata Fruhstorfer, 1911 : in Seitz, Grossschmett. Erde 9:381. TL. "Sikkim ; Assam ; Nepal". (NHML)
Elymnias malelas ivena Fruhstorfer, 1911 : in Seitz, Grossschmett. Erde 9:381. TL. "Tonkin ; Siam" [N.Vietnam ; Thailand]. (NHML)
Elymnias malelas nilamba Fruhstorfer, 1911 : in Seitz, Grossschmett. Erde 9:381. TL. nilamba, "Kumaon-Himalaya" [NW.India]. (NHML)


Type Materials Information

malelas ; ♂, Type (red), Rh5070, Sylhet. (NHML, examined)
subdecorata ; ♂, Type (red), Khasia Hill, Assam. (NHML, examined)
ivena ; ♂, Type (red), Aug.-Sep., Chiem-Hoa, "C. Tonkin" [N. Vietnam]. (NHML, examined) ; ♀, Type (red), Jan., "Hinlap, Siam" [Hin Lap, Saraburi, Thailand]. (NHML, examined)


Records taken from Literature

[ THAILAND ]
Elymnias malelas ivena ; Fruhstorfer,1911 : 381. (Siam)
Elymnias malelas ivena ; Godfrey,1916 : 123. (Bangkok; Pak Jong)
Elymnias malelas ivena ; Godfrey,1927 : 122. (N)
Elymnias malelas ivena ; Godfrey,1930 : 256. (Pak Jong; Bangkok)
Elymnias malelas ivena ; Pinratana,1988 : 9, pl.7, figs.10,♂,♂(Un),♀. (Chiang Mai; Nakhon Ratchasima; Loei; Chanthaburi; Nakhon Nayok)
Elymnias malelas ivena ; Yamaguchi & Aoki,2000 : 253, fig.70,♀. (Mukdahan: Phu Pha Taep)
Elymnias malelas ivena ; Katsuyama et al.,2005 : 67,68, fig.112,♂. (Umphang)
Elymnias malelas ivena ; Katsuyama et al.,2006 : 48, fig.84,♂. (Nan: Ban Nam Poon)
Elymnias malelas ; Ek-Amnuay,[2007] : 234, pl.89, figs.S10,♂,♂(Un),♀,♀(Un). (Doi Pha Hom Pok)
Elymnias malelas ; Les Day,2011-2022 : Samui Butterflies. (Nakhon Si Thammarat: Khao Luang) (accessed: 2022.01.01)
Elymnias malelas ; Ek-Amnuay,2012 : 232, pl.88, figs.N46,♂,♂(Un) ; pl.89, fig.N46,♀,♀(Un). (Doi Pha Hom Pok)
Elymnias malelas ivena ; Aoyama,2012 : 56. (Surat Thani: Kanchanadit)
Elymnias malelas ivena ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Elymnias malelas ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Chiang Mai; Phetchabun; Saraburi) (accessed: 2023.06.30)
Elymnias malelas ivena ; Kimura et al.,2016 : 164, figs.♂,♂(Un),♀,♀(Un). (Doi Par Hom Pok; Chiang Dao; Doi Suthep; Khun Lao Villege; Bang Salok; Phu Khiewo; Nakohn Nayok; Wan Nam Khiewo; Kratching WF; Pliewn WF; Kao Chamaoh; Kao Khiewo)
Elymnias malelas ; Wei et al.,2017 ; 104, fig.14, D♀ (Chiang Mai)
Elymnias malelas ; Küppers & Janikorn,2017 : 259, figs.Abb.S-070,♂(Un),S-071,♀(Un),S-189,S-197 (Phang Nga)
Elymnias malelas malelas ; Saito & Inayoshi,2018 : 29, figs.10,♂,♂(Un). (Yala)

[ LAOS ]
Elymnias malemas [sic] f. ivena ; Dubois & Vitalis,1924 : 31. (Vientiane)
Elymnias malelas ivena ; Motono & Negishi,1989 : 63, col.pl.14,♂ ; pl.81, figs.7,♂,8,♂(Un) ; pl.82, figs.1,♀,2,♀(Un). (Vientiane; Nam Moh; Hat Pho)
Elymnias malelas ivena ; Osada, Uémura & Uehara,1999 : 210, pl.87, figs.♂,♂(Un),♀,♀(Un). (Nong Het; Nhahin; Xiang Khouang; Xam Neua)
Elymnias malelas ivena ; Uémura,2003 : 2, fig.7,map.
Elymnias malelas ivena ; Uémura,2006 : 2, fig.5,map. (Phong Si)
Elymnias malelas ; Nakamura & Wakahara,2012 : 48.
Elymnias malelas malelas ; Onodera,2022 : 53, pl.157, figs.♂,♂(Un),♀,♀(Un). (Xiengkhouang: Tha; Phongsali)

[ CAMBODIA ]
Elymnias malelas ; Onodera,2007 : 22,24, fig.35♂. (Rattanakiri: Ta Veaeng; Yeak Laom)
Elymnias malelas ivena ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)
Elymnias malelas ; Chartier & Kosterin,2022 : 115. (Pursat; Ratanakiri)

[ VIETNAM ]
Elymnias malelas ivena ; Fruhstorfer,1911 : 381. (Tonkin)
Elymnias ivena ; Vitalis,1919 : 225. (Tonkin)
Elymnias malemas [sic] f. ivena ; Dubois & Vitalis,1924 : 31. (Tonkin)
Elymnias malelas ivena ; Metaye,1957 : 104. (N)
Elynmias malelas ivena ; Luong et al.,2004 : 3, 44, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Elymnias malelas ivena ; Monastyrskii,2005 : 32, pl.3, figs.3a♂,3b♂(Un),3c♀. (N: Muong Nhe; Ba Be; Na Hang; Huu Lien / C: Pu Hoat; Pu Mat; Ngoc Linh; Kon Cha Rang)
Elymnias malelas ivena ; Miyazaki, Saito & Saito,2006c : 28, fig.S-7,♀. (Lam Dong: Dambri; Dai Duc Me)
Elymnias malelas ; Miyazaki, Saito & Saito,2007c : 39, fig.S-7,♂. (Lam Dong: Dambri; Madagui)
Elymnias malelas ivena ; Monastyrskii & Devyatkin,2015 : 20. (N / C)
Elymnias malelas ; Wei et al.,2017 ; 104, fig.14, C♂ (Vietnam)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 4♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Rai : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 9♂ 3♀ (incl. photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 3♂ 2♀ (incl. photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 4♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 5♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Surat Thani : Kanchanadit dist. 1♂, 2010.08.19. Y.Harada (KS)
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Si Thammarat : Ron Phibun dist. 1♂, 2020.08.08 (photo by A.Giudici).
JanFebMarAprMayJun JulAugSepOctNovDec

Yala : 1♂ 1♀ ; Betong dist.
1♀, 2017.01.12. (photo by K.Yano & Y.Inayoshi) ; 1♂, 2017.03.21. E.Ikari (EI).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Xieng Khouang : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Luang Prabang : 1♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Oudomxai : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Vientiane : 3♂ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Vinh Phuc : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Lam Dong : 2♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes

The butterfly is locally not rare in montane forests at low to moderate elevations (Alt. 300 - 1600m). It is a good mimic of Euploea mulciber (Danainae).
This species was known for a long time only from the continental part, however it has recently been recorded from peninsular Thailand and West Malaysia, as mentioned in Kimura et al.,2016. The distribution range of this species is expanding rapidly.


Literature

Doubleday, E.,1844-1848 : List of the Specimens of lepidopterous Insects in the Collection of the British Museum.
Part 1, 2 , and Appendix. London. British Museum 12 mo 1:vi + 150pp; 2:ii+57pp; Appx:ii+38pp.

Hewitson, W.C.,1852-1877 : Illustrations of new species of exotic Butterflies, selected chiefly from the Collections of W.Wilson Saunders and William C. Hewitson.
5 vols. John Van Voorst, London.

Fruhstorfer, H.,1911 : 4. Familie: Satyridae in Seitz, Gross-Schmett. Erde 9
377-384. (20.Jun.1911)

Aoyama, Y.,2012 : A first record of Elymnias malelas ivena Fruhstorfer,1911 in Malaysia.
Butteflies (Teinopalpus) 61:56. (in Japanese)

Wei, C.H., Lohman, D., Peggie, D. & Yen, S.H.,2017 : An illustrated checklist of the genus Elymnias Hübner,1818 (Nymphalidae, Satyrinae).
Zookeys 676:47-152.


Elymnias malelas ♂ Elymnias malelas ♀
♂, Chiang Mai, Thailand. ♀, Lampang, Thailand.
Elymnias malelas ♂ Elymnias malelas ♀
♂, Chiang Mai, Thailand. ♀, 2017.01.12. Yala, Thailand. (photo by K.Yano)
Elymnias malelas ♂
♂, Nakhon Si Thammarat, Thailand. (photo by A.Giudici)

1998.12.09 - 2023.07.31.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © Y.Inayoshi.