♂, Vinh Phuc, Vietnam. (photo by K.Saito)		♀, Vinh Phuc, Vietnam. (photo by K.Saito)
♂, Lao Cai, Vietnam. (photo by K.Saito)		♀, Lao Cai, Vietnam. (photo by K.Saito)
♂, Chanthaburi, Thailand.		♀, Lao Cai, Vietnam. (photo by K.Saito)

ON : Delias acalis shinkaii Morita,1998 OD : Wallace 4(2):28, pl.12, figs.9(HT,♂), 10(AT,♀). TL : Tam Dao, N.Vietnam. (in coll. S.Morita) Distribution : SE.Thailand, Cambodia, Vietnam. ● ssp.pyramus ● ssp.shinkaii ● ssp.perakana

Holotype ♂, 24.Aug.1997. Mt.Tam Dao, 1200m, N.Vietnam. A.Shinkai leg. (in coll. S.Morita)
Paratypes : 23♂ 14♀, same locality as holotype.

[ THAILAND ]
Delias thysbe thysbe ; Davidson & Macbeth,1938 : 70. (Chanthaburi)

[ CAMBODIA ]
Delias acalis acalis ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)
Delias acalis shinkaii ; Kosterin,2019b : 26. (Mondulkiri)
Delias acalis shinkaii ; Chartier & Kosterin,2022 : 114. (Mondulkiri; Pursat)

[ VIETNAM ]
Delias pyramus ; Joannis,1901 : 328. (Tonkin: Cao Bang)
Delias thysbe ; Vitalis,1919 : 217. (Annam)
Delias thysbe f. thysbe ; Dubois & Vitalis,1921 : 18. (Tonkin)
Delias thysbe f. pyramus ; Dubois & Vitalis,1921 : 18. (Annam)
Delias thysbe pyramus ; Lemée,1950 : 5. (Hagiang)
Delias thysbe thysbe ; Metaye,1957 : 85. (N)
Delias thysbe pyramus ; Metaye,1957 : 85. (N / C)
Delias acalis ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Bach Ma)
Delias acalis shinkaii ; Miyazaki, Saito & Saito,2006 : 10, figs.Pi-2,♂,♀. (Lam Dong: Dambri)
Delias acalis pyramus ; Monastyrskii & Devyatkin,2015 : 15. (N / C: Lam Dong)
Delias acalis shinkaii ; Yokochi et al.,2015 : 39, figs.15,♂,15v♂(Un),16,♀,16v♀(Un). (Khanh Hoa: Hon Ba)

[ THAILAND ]
Chanthaburi : Soi Dao dist. 1985.02.12. 1♂, P.Sukkit (YI).

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[ VIETNAM ]
Lao Cai : 1♂ 2♀.

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Vinh Phuc : 2♂ 1♀.

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Lam Dong : 2♂ 2♀.

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This taxon is intermediate between ssp.pyramus and the nominotypical subspecies. Some examples from Lao Cai, N.W.Vietnam tend to have closer markings to ssp.pyramus.
The populations in S.Vietnam and S.E.Thailand seem to be slightly different from ssp.shinkaii. Monastyrskii et al.,2011 recorded it from Cambodia and treated it as the nominotypical subspecies. I consider this population to be identical to S.E.Thailand - S.Vietnam ones from a biogeographic point of view. Further taxonomic study is needed.

Morita, S.,1998 : A new subspecies of Delias acalis Godart,1819 from Vietnam (Lepidoptera: Pieridae)
Wallace 4(2):27-28, pl.12.

1999.04.22 - 2024.03.31.