Papilio memnon agenor   Linnaeus,1758

Papilio memnon agenor ♂ Up. Papilio memnon agenor ♂ Un. Papilio memnon agenor ♀ Up. Papilio memnon agenor ♀ Un.
♂, 1995.03.09. Doi Suthep, Chiang Mai, Thailand. ♀, 1991.03.05. Wiang Pa Pao, Chiang Rai, Thailand.
Papilio memnon agenor ♂ Up. Papilio memnon agenor ♂ Up. Papilio memnon agenor ♀ Up. Papilio memnon agenor ♀ Up.
♂, 1991.03.--. Doi Phu Kha, Nan, Thailand. ♂, 1991.06.--. Doi Phu Kha, Nan, Thailand. ♀, 1990.01.--. Chiang Dao, Chiang Mai, Thailand. ♀, 1991.04.01. Doi Phu Kha, Nan, Thailand.
Papilio memnon agenor ♀ Up. Papilio memnon agenor ♀ Up. Papilio memnon agenor ♀ Up. Papilio memnon agenor ♀ Un.
♀, 1990.01.--. Chiang Dao, Chiang Mai, Thailand. ♀, 1992.03.20. Than To, Yala, Thailand. ♀, 1994.03.30. Than To, Yala, Thailand.

ON : Papilio agenor  Linnaeus,1758
OD : Syst. nat. (ed. 10) 1:460.
TL : "Asia".



Distribution :
Nepal, India to Myanmar, Thailand, Laos, Cambodia, Vietnam, China,
Hainan, Langkawi, W.Malaysia, Tioman, Singapore.
Papilio memnon agenor Map

Synonyms

Papilio androgeos  Cramer,[1776] : Uitl. Kapellen 1(8):142, pl.91, figs.A,B.  TL. China.
Papilio alcanor  Cramer,[1777] : Uitl. Kapellen 2(9-16):107, pl.166, fig.A♀.  TL. China.
Papilio esperi  Butler,[1879] : Trans. linn. Soc. Lond. (2) Zool. 1:553, pl.68, fig.7.  TL. Malacca ; Penang.
Papilio phoenix  Distant,1885 : Rhop. malayana : 340, pl.27, fig.7(♀).  TL. Malay Pen.
Papilio cilix  Distant,1885 : Rhop. malayana 340-341, pl.29, fig.4(♂),5(♀).  TL. Malay Pen.
Papilio androgeos var.depelchini  Robbe,1892 : Annls. Soc. ent. Belg. 36:125.  TL. Kurseong, Sikkim.
Papilio memnon agenor ♂ab.primigenius  Rothschild,1895 : Novit. zool. 2(3):319.  TL. Sikkim ; Khasia Hills.
Papilio memnon agenor ♀ab.butlerianus  Rothschild,1895 : Novit. zool. 2(3):320. 
Papilio memnon agenor ♀ab.distantianus  Rothschild,1895 : Novit. zool. 2(3):320.  TL. India ; Burma ; Siam ; Malay Pen ; Hainan.
Iliades polymnestoroides  Moore,1902 : Lep. Ind. 5:202-203, pl.451, figs.1♂,2♀.  TL. Jaintia Hills, Assam.
Papilio memnon agenor ♀ab.vinius  Fruhstorfer,1902 : Deut. ent. Zeit. [Iris] 15:307.  TL. Malay Pen.
Papilio memnon agenor ♀f.rhetenorina  Jordan,1909 : in Seitz, Grossschmett. Erde. 9:72.  TL. N.India.
Papilio memnon agenor ♀f.phoeniciana  Strand,1916 : Lep. Niepelt. (2):25, pl.13, fig.8♀.  TL. Sikkim.
* Papilio memnon ♀f.aphrodite  Rober,1927 : Int. ent. Z. 21(13):97.  TL. Naga Hills.
Papilio memnon aurantiaca  Rousseau-Decelle,1933 : Bull. Soc. ent. Fr. 38(17):273.  TL. Tonkin ?.(♂) (homonym).
* Papilio ab.pohli  Dufrane,1946 : Bull. Annls Soc. ent. Belg. 82:116.
* Papilio f.subvenusia  Dufrane,1946 : Bull. Annls Soc. ent. Belg. 82:116.
Papilio memnon f.indiv.♂coeruleocinetus  Rousseau-Decelle,1947 : Bull. Soc. ent. Fr. 51:129.  TL. Hoa Binh, Tonkin.(♂)
Papilio memnon agenor ♀f.tanahsahi  Eliot,1982 : Malay Nat. J. 35:179.  TL. Perak, Peninsular Malaysia. (♀,BMNH)


Type Materials Information

esperi ; ♀, Type (red), Rh.11024, Malacca. Pinwell. (BMNH, examined.)
phoeniciana ; ♀, Type HT (red), Sikkim. Strand det. (BMNH, examined.)
tanahsahi ; ♀, Holotype, 1975.ii.22, Ulu Groh 800 ft, Gopeng, Perak, Peninsular Malaysia. (BMNH)


Records taken from Literature

[ THAILAND ]
Papilio memnon agenor ; Fruhstorfer,[1903] : 174. (Siam)
Papilio memnon agenor ; Godfrey,1916 : 110.
Papilio memnon agenor ; Pendlebury,1923 : 24. (Nakhon Sri Thammarat: Tai Sai River ; Khao Ram)
Papilio memnon agenor ; Godfrey,1927 : 120. (North)
Papilio memnon agenor ; Godfrey,1930 : 213-214. (all over)
Papilio memnon agenor ; Godfrey,1932a : 264. (Peninsula)
Papilio memnon ♀f.alcanor ; Davidson & Macbeth,1938 : 68. (Lampang: Me Htam Forest / Paknampoh)
Papilio memnon agenor ; Reeves,1966 : 10. (Nakhon Nayok: Khao Yai)
Papilio memnon agenor f.distantianus ; Reeves,1966 : 10. (Nakhon Nayok: Khao Yai)
Papilio memnon agenor f.distantianus ; Reeves,1967a : 129. (Nakhon Nayok)
Papilio memnon agenor ; Pinratana,1977 : 41, fig.P29,♂. (all over)
Papilio memnon agenor ♀f.distantianus ; Pinratana,1977 : 41, fig.P29,♀. (all over)
Papilio memnon agenor ♀f.esperi ; Pinratana,1977 : 42, fig.P29,♀. (Yala)
Papilio memnon agenor ♀f.butlerianus ; Pinratana,1977 : 42, fig.P29,♀. (Trang / Yala)
Papilio memnon agenor ♀f.agenor ; Pinratana,1977 : 43, fig.P29,♀. (all over)
Papilio memnon agenor f.agenor ; Pinratana,1992 : 24-25,43, pl.39 fig.31,♂,♂(variation) ; pl.40, fig.31,♀. (Chiang Mai / Nakhon Nayok / Chanthaburi / Kanchanaburi / Bangkok / Ratchaburi / Trang)
Papilio memnon agenor f.distantianus ; Pinratana,1992 : 24-25,43, pl.40, fig.31,♀. (Nakhon Nayok / Ratchaburi / Ranong / Trang)
Papilio memnon agenor f.esperi ; Pinratana,1992 : 24-25,43, not fig. (List up only.)
Papilio memnon agenor f.butlerianus ; Pinratana,1992 : 24-25,43, pl.41, fig.31,♀. (Ranong / Yala)
Papilio memnon agenor f.thetenorina ; Pinratana,1992 : 24-25, pl.41, fig.31,♀. (No Data)
Papilio memnon agenor f.vinius ; Pinratana,1992 : 24-25, pl.42, fig.31,♀. (No Data)
Papilio memnon agenor ; Pinratana,1992 : 24-25, pl.42, fig.31,Gynandromorph.
Papilio memnon agenor ; Yamaguchi & Aoki,2000 : 251, fig.41♀. (Mukdahan: Phu Mu ; Phu Shang)
Papilio memnon agenor ; Yamaguchi & Aoki,2002 : 16. (Mukdahan: Ban Lao Kram ; Phu Huay Sing Ban Na Lak ; Ban Gok Tum / Ubon Ratchathani: Kaeng E-Keow)
Papilio memnon agenor ; Yamaguchi, Aoki & Akiyama,2005 : 45-46. (Ubon Ratchathani: Kaen Tana / Mukdahan: Phu Hin Khan ; Nam Tok Tad Ton)
Papilio memnon agenor ; Katsuyama et al.,2005 : 64, fig.42♂. (Umphang)
Papilio memnon agenor ; Katsuyama et al.,2006 : 46, fig.29♂. (Nan: Ban Nam Poon)
Papilio memnon agenor ; Ek-Amnuay,[2007] : 84, pl.13, fig.P30,♂,♂(Un),♂♀(Gynandromorph) ; pl.14, fig.30,♀(f.agenor), ♀(f.agenor),♀(f.distantianus),♀(f.distantianus),♀(f.alcanor),♀(f.rhetenorina),♀(f.butlerianus),♀(f.esperi). (Chiang Mai / Chanthaburi / Ranong / Yala)
Papilio memnon agenor ; Les Day,2011- : Samui Butterflies, fig.♂,♂(Un),♀f.agenor,♀f.butlerianus,♀f.distantianus,♀f.esperi(Un). (Koh Samui)
Papilio memnon agenor ; Kimura et al.,2011 : 126, fig.♂,♂(Un); f.butreianus♀,♀(Un); f.vinus♀,♀(Un); f.agenor♀,♀(Un); f.distantianus♀,♀(Un). (Maethro / Bang Salok / Thung Saleng Luang / Phu Khiewo / Khao Yai / Sarika / Nakohn Nayork / Wan Nam Khiewo / Kao Soi Dao / Kratchin WF / Khao Khiewo / Khao Phang WF / Sai York Noi / Kuawai Yai / Khlong Naka / Kam Puang / Ka Chong / Satun / Thaleban / Than To WF)
Papilio memnon agenor ; Ek-Amnuay,2012 : 84, pl.13, fig.P28,♂,♂(Un),♂♀(Gynandromorph) ; pl.14, fig.28,♀(f.agenor), ♀(f.agenor),♀(f.distantianus),♀(f.distantianus),♀(f.alcanor),♀(f.rhetenorina),♀(f.butlerianus),♀(f.esperi). (Chiang Mai / Chanthaburi / Ranong / Yala)

[ LAOS ]
Papilio memnon agenor ; Motono & Negishi,1989 : 46, pl.6, figs.3♂,4♂(Un),5♀,6♀(Un),7♀,8♀(Un). (Luang Prabang / Vientiane / Simouang / Tha Ngone / Ban Houei Sai / Nam Moh / Nam Ngum Dam)
Papilio memnon agenor ; Osada, Uémura & Uehara,1999 : 200, pl.14, figs.♂,♂(Un) ; pl.15, figs.♀(f.agenor),♀(f.distantianus). (Thabok / Luang Phrabang / Muang Khouang)
Papilio memnon agenor ; Nimura & Wakahara,2005 : 38-39. (Namdhua / Thakhek)
Papilio memnon agenor ; Cotton & Racheli,[2007] : 359-361. (throughout Laos)
Papilio memnon agenor ; Onodera,2009b : 34-35. (Vientinane: Namsong Riverside)
Papilio memnon agenor ; Nakamura & Wakahara,2012 ; 41.

[ CAMBODIA ]
Papilio memnon agenor ; Woodfield & Murton,2006 : figs.♂,♂(Un),♀,♀(Un). (Botum Sakor NP.)
Papilio memnon ; Onodera,2007 : 26,27, fig.7♂. (Rattanakiri: Ou Chum / Siem Reap: Phnum Kuhlen)
Papilio memnon agenor ; Monastyrskii et al.,2011 : 128. (Phnom Samkos W.S.)

[ VIETNAM ]
Papilio agenor ; Joannis,1901 : 333. (Tonkin: Cao Bang)
Papilio memnon agenor ; Fruhstorfer,[1903] : 174. (Tonkin: Chime-Hoa / Annam: Xom-Gom; Phuc-Son)
Papilio agenor ; Lemee,1950 : 3. (Hagiang / Langson)
Papilio memnon f.♀.alcanor ; Dubois & Vitalis de Salvaza,1914 : 149. (N.Tonkin)
Papilio memnon agenor ; Metaye,1957 : 83. (North / Central / South)
Papilio memnon agenor f.♀.agenor ; Metaye,1957 : 83. (North)
Papilio memnon agenor f.♀.alcanor ; Metaye,1957 : 83. (North / Central / South)
Papilio memnon agenor f.♀.distantianus ; Metaye,1957 : 83. (North / Central / South)
Papilio memnon agenor ; Ikeda et al.,1998 : 17, figs.4:6♂,7♂(Un),8♀, 9:9(♂genitalia). (Cuc Phuong)
Papilio memnon agenor ; Shinkai,1999 : 55. (North: Tam Dao / South: Bao Lac)
Papilio memnon agenor ; Miyazaki, Saito & Saito,2006 : 7, figs.Pa-18♂,♀,♀. (Lam Dong: Dambri ; Gung Re ; Bao Loc ; Dai Duc Me ; Madagui / Dong Nai: Thac Mai / Bin Thuan: Dong Tien ; Ta Cu / Ho Chi Minh)
Papilio memnon ; Monastyrskii et al.,2006 : 38. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Papilio memnon agenor ; Monastyrskii,2007 : 61-62, pl.16, figs.a♂,b♀(f.distantianus),c♀(f.alcanor),d♀(f.agenor),e♀(f.agenor),f♀(f.agenor). (throughout Vietnam)


Specimens Examined

[ THAILAND ]
Mae Hong Son, Ban Na Plachat.
1984.03.11
1♂


Y.Inayoshi (YI)
Chiang Rai, Mae Kajan.
1987.02.--


2♀
Local Catcher (YI)
Chiang Rai, Wiang Pa Pao.
1991.03.05


1♀
M.Nishimura (MN)
Chiang Mai, Chiang Dao.
1990.01.--


2♀
M.Nishimura (MN)
Chiang Mai, Chiang Dao.
1987.04.01


1♀
Local Catcher (YI)
Chiang Mai, Doi Suthep.
1987.02.01
1♂


Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1984.02.05
1♂


Y.Shiraishi (YI)
Chiang Mai, Doi Suthep.
1983.03.01
1♂


Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1983.03.02
1♂
1♀
Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1983.03.05


1♀
Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1995.03.09
1♂


M.Nishimura (MN)
Chiang Mai, Doi Suthep.
1987.04.02


1♀
Y.Inayoshi (YI)
Chiang Mai, Doi Suthep.
1987.04.06
1♂


Y.Inayoshi (YI)
Phrae, Wang Chin.
1989.10.16


1♀
Local Catcher (YI)
Lampang, Jae Hom.
1988.03.04


1♀
Local Catcher (YI)
Lampang, Mae Hom.
1989.10.31


1♀
Local Catcher (YI)
Nan, Doi Phu Kha.
1991.03.--
2♂
1♀
Local Catcher (YI)
Nan, Doi Phu Kha.
1991.04.--


1♀
Local Catcher (YI)
Nan, Doi Phu Kha.
1991.06.--
1♂
1♀
Local Catcher (YI)
Tak, Umphang.
1994.03.19
1♂


Y.Inayoshi (YI)
Trat, Koh Chang.
1996.03.18
1♂


Y.Inayoshi (YI)
Ranong, Ranong.
2012.03.03
2♂
1♀
E.Ikari (EI)
Ranong, Khlong Naka.
1983.03.21
1♂


Y.Inayoshi (YI)
Ranong, Khlong Naka.
1983.04.12
1♂


Y.Inayoshi (YI)
Ranong, Muang Chone.
1994.11.28
1♂


S.Ruangrit (YI)
Nakhon Si Thammarat, Ka Rom WF.
1992.03.12
1♂


Y.Inayoshi (YI)
Satun, Koh Tarutao.
1987.03.17
1♂


Y.Inayoshi (YI)
Yala, Than To.
1992.03.20


1♀
Y.Inayoshi (YI)
Yala, Than To.
1987.03.21
1♂


Y.Inayoshi (YI)
Yala, Than To.
1987.03.22
1♂


Y.Inayoshi (YI)
Yala, Than To.
1987.03.24
1♂


Y.Inayoshi (YI)
Yala, Than To.
1994.03.30


1♀
Y.Inayoshi (YI)

[ LAOS ]
Xiang Khouang, Muang Phon Savan.
1996.06.26
1♂


P.Sukkit (YI)
Lak Sao, Nam Phao.
2005.02.26-03.11
1♂


A.M.Cotton (AC)
Lak Sao, Nam Phao.
2005.03.21-03.30
2♂


A.M.Cotton (AC)
Vientiane, Phu Khao Kway.
1996.04.01
2♂


Y.Inayoshi (YI)
Thabok, Tad Leuk.
2005.02.26-03.11
10♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.02.26-03.11


1♀
A.M.Cotton (AC)
Thabok, Tad Xai.
2005.06.14
1♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.06.23
3♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.06.29
1♂


A.M.Cotton (AC)
Thabok, Tad Xai.
2005.08.08-08.10
32♂


A.M.Cotton (AC)
Hinboun, Koun Kham.
2005.02.26-03.11
2♂


A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.06.15


1♀
A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.06.17
1♂


A.M.Cotton (AC)
Hinboun, Koun Ngun.
2005.07.30-08.07
16♂
20♀
A.M.Cotton (AC)
Hinboun,Phu Pha Man.
2005.08.01-08.07


13♀
A.M.Cotton (AC)

[ VIETNAM ]
Lam Dong, Dambri.
2003.09.17


1♀
K.Saito (KS)
Lam Dong, Dambri.
2003.09.20


1♀
K.Saito (KS)
Lam Dong, Dai Duc Me.
2003.12.29
1♂


K.Saito (KS)


Notes
Literature

Linnaeus, C.,1758 : Systema naturae per regna tria naturae,secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis.
Holmiae, Laurentil Salvii. (Edition 10). 1(Animalia):[iv]+824pp.

Cramer, P.,[1775]-[1776] :
Uitlandsche Kapellen. voorkomende in de drie Waereld-Deelen Asia, Africa en America, by een verzameld en bescreeven.
Amsterdam. Baalde, and Utrecht, Wild. 1(1-7):xxx + 16 + 132pp, pl.1-84, [1775] ; 1(8):133-156, pl.85-96,[1776].

Cramer, P.,[1777] :
Uitlandsche Kapellen. voorkomende in de drie Waereld-Deelen Asia, Africa en America, by een verzameld en bescreeven.
Amsterdam. Baalde, and Utrecht, Wild. 2(9-16):1-152, pl.97-192.

Butler, A.G.,[1879] : The Butterflies of Malacca.
Trans. linn. Soc. Lond., (2)Zool. 1(8):533-568, pl.68,69.

Distant, W.L.,1882-1886 : Rhopalocera Malayana : a Description of the Butterflies of the Malay Peninsula.
London, Author, Penang, Logan : xvi+482+[4]pp, 46 pls.

Robbe, H.,1892 : Insectes du Bengale occidental.
Liste d'une collection de Lepidopteres recueillis au Bengale occidential; avec une description d'une variete nouvelle et quelques considerations sur des especes connues.
Annls Soc. ent. Belg. 36:122-131.

Rothschild, L.W.,1895 : A revision of the Papilios of the Eastern Hemisphere, exclusive of Africa.
Novit. zool. 2(3):167-463, pl.4.

Moore, F.,1901-1902 : Lepidoptera Indica. vol.V. Rhopalocera, Family Nymphalidae, Riodinidae, Papilionidae.
5:1-96, 1901; 97-225, 1902. L.Reeve & Co., London.

Fruhstorfer, H.,1902 : Neue Papilioformen und andere Lepidopteren aus Ost-Asien und dem malayischen Archipel.
Deut. ent. Zeit. [Iris] 15:306-315.

Strand, E.,1914-1916 : Lepidoptera Niepeltiana.
Abbidungen und Beschreibungen neuer und wenig bekannter Lepidoptera aus der Sammlung W.Niepelt.
Leipzig, Druck von Urban 4to (1):1-65, pls.1-12(1914); (2):[4]+1-27pp, pls.13-17(1916).

Rousseau-Decelle, G.,1933 : Notes sur queques formes nouvelles des genres Papilio et Charaxes. (Lep.)
Bull. Soc. ent. Fr. 38(17):269-273.

Rousseau-Decelle, G.,1947 : Contribution a letude des Papilio de la faune Indo-Oceanienne (Lep. Papilionidae)
Bull. Soc. ent. Fr. 51:128-133.

Eliot, J.N.,1982 : On three swallowtail butterflies from Peninsular Malaysia.
Malay Nat. J. 35:179-182.

Papilio memnon agenor ♀
♀, Laos. ( photo by K.Phongpiyamit )

1996.10.02 - 2014.06.14.