Famegana nisa  (Wallace,1866)

Famegana nisa ♂ Up. Famegana nisa ♂ Un. Famegana nisa ♀ Up. Famegana nisa ♀ Un.
♂, Phetchaburi, Thailand. (photo by K.Yano) ♀, Phetchaburi, Thailand. (photo by K.Yano)
Famegana nisa ♂ Up. Famegana nisa ♂ Un. Famegana nisa ♀ Up. Famegana nisa ♀ Un.
♂, Surat Thani (Koh Tao), Thailand. (photo by P.Kosierb) ♀, Surat Thani (Koh Tao), Thailand. (photo by P.Kosierb)
Famegana nisa ♂ Up. Famegana nisa ♂ Un. Famegana nisa ♂ Up. Famegana nisa ♂ Un.
♂, Xiang Khouang, Laos. (photo by K.Saito) ♂, Xiang Khouang, Laos. (photo by K.Saito)

ON : Lycaena nisa Wallace,1866
OD : Proc. zool. Soc. Lond. 1866:360-361.
TL : "Takaw, Formosa" (♀, NHML).



Distribution : Thainand, Laos, Vietnam, S.China, Taiwan,
Philippines, Sulawesi, Australia, Vanuatu, Fiji, Samoa, Tonga.
alsulus eggletoni map

Synonyms

Lycaena alsulus Herrich-Schaffer,1869 : Stettin ent. Ztg. 30(1-3):75. TL. Rockhampton and Upolu [Australia].
Lycaena exilis Lucas,1889 : Proc. R. soc. Qd. 6(4):159-160, figs.13-15. TL. Cooktown to Bowen [Australia]. (Homonym)
Lycaena lulu Mathew,1889 : Trans. ent. Soc. Lond. 1889(2):312-314. TL. Tongatabu, [Tonga].
Lycaena gracilis Miskin,1890 : Proc. linn. Soc. N.S.Wales (2)5(1):37. TL. Brisbane to Cooktown [Australia].
* Lycaena exiloides Lucas,1891 : Proc. R. soc. Qd. 7(1):47. Replacement name for Lycaena exilis.
Zizera kalawarus Ribbe,1926 : Ent. Mitt. 15(1):91. TL. Celebes.
* Zizera taiwana Sonan,1938 : Zephyrus 7:254. TL. "Inrin, Formosa". (♂, NHML)
Zizeeria alsulus eggletoni Corbet,1941 : Hongkong Nat. 10(3/4) : 150-151. TL. Hong Kong. (♂, NHML)


Type Materials Information

nisa ; Lectotype ♀, Formosa. designated by Hsu,2020. (NHML)
taiwana ; Holotype ♂, 1932.x.30. Inrin. (NHML) ; Allotype ♀, same data as holotype (NHML) ; Paratype 1♀, same data as holotype (NHML)
eggletoni ; Holotype ♂, 1914.ix.5. Hong Kong dist., New Territory. R.W.Barney (NHML) ; Paratype 1♂, same data as holotype (NHML)


Records taken from Literature

[ THAILAND ]
Femegana alsulus ; Ek-Amnuay,[2007] : 570, 718, fig.52. (Nakhon Si Thammarat / Samui)
Famegana alsulus eggletoni ; Les Day,2011 - : Butterflies of Koh Samui, fig. (Koh Samui / Koh Phangan) (accessed: 2021.01.30)
Femegana alsulus eggletoni ; Ek-Amnuay,2012 : 589, pl.267, figs.L128,♂,♂(Un),♀. (Nakhon Si Thammarat / Samui / Phetchaburi)
Femegana alsulus eggletoni ; Kimura et al.,2014 : 182. (by literature)

[ LAOS ]
Femegana alsulus eggletoni ; Osada, Uémura & Uehara,1999 : 215, pl.108, figs.♂,♀(Un). (Xiang Khouang)
Femegana alsulus eggletoni ; Nakamura & Wakahara,2012 : 52.

[ VIETNAM ]
Femegana alsulus ; Monastyrskii,2005b : 158. (Kon Tum: Ngoc Linh)
Femegana alsulus eggletoni ; Miyazaki et al.,2010 : 28, figs.L-163,♂,♀. (Phan Thiet)
Femegana alsulus ; Monastyrskii & Devyatkin,2015 : 52. (C: Ninh Thuan)
Femegana alsulus eggletoni ; Saito et al.,2017 ; 80. (S: Nui Coto)


Specimens Examined

[ THAILAND ]
Phetchaburi : 10♂ 3♀ (incl.photo by T.Aoyama).
JanFebMarAprMayJun JulAugSepOctNovDec

Surat Thani (Koh Phangan dist: Koh Phangan ; Koh Tao) : 3♂ 2♀ (incl.photo by A.Giudici).
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Xiang Khouang : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
This is very localized species. The butterfly is found in wasteland or open land at low elevations, especially is found in islands or sea side.

Literature

Wallace A.R. & F.Moore,1866 : List of Lepidopterous insects collected at Takaw, Formosa, by Mr.Robert Swinhoe
Proc. zool. Soc. Lond. 1866:355-365.

Herrich-Schaffer, G.A.W.,1869 : Neue Schmetterlinge aus dem "Museum Godeffroy" in Hamburg
Stettin ent. Ztg. 30(1-3):65-80, pls.1-4.

Lucas, F.H.,1889 : Six new species of Rhopalocera.
Proc. R. soc. Qd. 6(4):155-161.

Mathew, G.F.,1889 : Descriptions and life-histories of new species of Rhopalocera from the Western Pacific
Trans. ent. Soc. Lond. 1889(2):311-315.

Miskin, W.H.,1890 : Descriptions of hitherto undescribed Australian Lepidoptera (Rhopalocera), principally Lycaenidae
Proc. linn. Soc. N.S.Wales (2)5(1):29-43.

Ribbe, C.,1926 : Neue Lycaenenformen hauptsachlich von Celebes (Lep.: Lycaenidae).
Ent. Mitt. 15(1):78-91.

Corbet, A.S.,1941 : A List of the Butterflies of Hong Kong (3).
Hongkong Nat. 10(3/4):148-165A, 166, 5 figs, 1 pl.

Hsu, Y-F.,2020 : The identity of Alfred Wallace's mysterious butterfly taxon Lycaena nisa solved: Famegana nisa comb. nov., a senior synonym of F.alsulus (Lepidoptera, Lycaenidae, Polyommatinae)
ZooKeys 966:153-162.

Famegana nisa ♂ Famegana nisa ♂
♂, Phetchaburi, Thailand. (photo by T.Aoyama) ♂, Phetchaburi, Thailand. (photo by T.Aoyama)
Famegana nisa ♂ Famegana nisa
♂, Surat Thani (Koh Phangan), Thailand. (photo by A.Giudici) Surat Thani (Koh Phangan), Thailand. (photo by A.Giudici)
Famegana nisa ♂ Famegana nisa ♂
♀+♂, Surat Thani (Koh Phangan), Thailand. (photo by A.Giudici) ♂, Surat Thani (Koh Phangan), Thailand. (photo by A.Giudici)

2009.03.10 - 2021.01.31.
A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © 1996- Y.Inayoshi.