♂ (dsf), Chiang Mai, Thailand.		♀ (dsf), Nan, Thailand.
♂ (wsf), Nan, Thailand.		♀ (wsf), Chiang Mai, Thailand.
♂ (dsf), Phetchaburi, Thailand.		♂ (dsf), Phetchaburi, Thailand.
♂, Chiang Mai, Thailand.		♂ (ab.), Chiang Mai, Thailand.

ON : Cyaniris puspa gisca Fruhstorfer, 1910 OD : Stett. ent. Ztg. 71(2):285. TL : 'Sikkim; Assam; Birma' [Sikkim, N.India]. (NHML) Distribution : NE.Afghanistan to Myanmar, Thailand, Cambodia, Laos, Vietnam, S.China. ● ssp.gisca ● ssp.lambi

Cyaniris puspa hermagoras Fruhstorfer, 1910 : Stett. ent. Ztg. 71(2):284. TL. Hainan. (NHML)
Lycaenopsis puspa gisca forma artena Fruhstorfer, 1917 : Arch. Naturgesch. (A)82(1):25. TL. 'Sikkim ; Assam ; Oberbirma' [Sikkim, N.India]. (NHML) (IFS)
Lycaenopsis puspa telis Fruhstorfer, 1917 : Arch. Naturgesch. (A)82(1):25. TL. Andaman. (NHML)
Celastrina puspa barneyi Corbet, 1941 : Hongkong Nat. 10(3/4):149. TL. Hong Kong. (NHML)

gisca ; Lectotype ♂, Sikkim, India. designated by Eliot & Kawazoé,1983. (NHML)
hermagoras ; Lectotype ♀, Hainan. designated by Eliot & Kawazoé,1983. (NHML)
artena ; Lectotype ♂, Sikkim. designated by Eliot & Kawazoé,1983. (NHML)
telis ; Lectotype ♂, Andamans. designated by Eliot & Kawazoé,1983. (NHML)
barneyi ; Holotype ♂, Hong Kong. (NHML)

[ THAILAND ]
Cyaniris puspa imperatrix ; Godfrey,1916 : 135. (Chonburi: Dong Rek Hills)
Lycaenopsis puspa gisca ; Godfrey,1927 : 125. (N)
Lycaenopsis puspa gisca ; Godfrey,1930 : 317. (Doi Chang; Prae: Me Song forest; Pak Jong; Muak Lek; Chonburi: Sriracha)
Acytolepis puspa gisca ; Pinratana,1981 : 44, 162, pl.7, figs.1,♂,2,♂,3,♂(Un) ; pl.34, fig.3,♂(dsf) ; pl.35, fig.2,♂(dsf,Un). (Chiang Mai; Chaiyaphum; Chonburi; Nakhon Nayok; Ratchaburi)
Acytolepis puspa gisca ; Yamaguchi, Aoki & Akiyama,2005 : 53, fig.25,♂. (Ubon Ratchathani: Phu Pung)
Acytolepis puspa gisca ; Katsuyama et al.,2005 : 66, fig.71,♂. (Umphang: Tha Sai; Umphang)
Acytolepis puspa gisca ; Katsuyama et al.,2006 : 47, fig.52,♂. (Nan: Ban Nam Poon)
Acytolepis puspa gisca ; Ek-Amnuay,[2007] : 564, pl.255, figs.L88a,♂,♂(Un),♀[♂],♀[♂](Un). (Wiang Papao; Wang Chin)
Acytolepis puspa gisca ; Ek-Amnuay,2012 : 582, pl.264, figs.L113a,♂,♂(Un),♀. (Wiang Pa Pao; Wang Chin; Nakhon Nayok)
Acytolepis puspa gisca ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Acytolepis puspa gisca ; Kimura et al.,2014 : 187, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Doi Pha Hom Pok; Doi Ang Khaang; Chiang Dao; Doi Suthep; Doi Inthanon; Thung Saleng Luang; Phu Kradung; Phu Khiewo; Kao Yai; Nakohn Nayok; Wan Nam Khiewo; Kao Soi Dao; Kratchin WF; Kao Saverb; Kao Khiewo; Doi Musso; Thung Yai; Sai Yok Noi; Kaeng Kratchan)
Celatoxia marginata splendens ; Kimura et al.,2014 : 188, figs.♀,♀(Un). (Kao Soi Dao)
Acytolepis puspa gisca ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Chanthaburi; Ratchaburi) (accessed: 2023.03.11)

[ LAOS ]
Acytolepis puspa gisca ; Motono & Negishi,1989 : 69, pl.99, figs.B5,♂,B6,♂(Un),B7,♀,B8,♀(Un). (Luang Prabang; Vientiane; Pakse; Vang Vieng)
Acytolepis puspa gisca ; Osada, Uémura & Uehara,1999 : 214, pl.109, figs.♂,♂(Un),♀,♀. (Oudomxay; Xaignabouri; Lak Sao)
Acytolepis puspa gisca ; Nimura & Wakahara,2005 : 46. (Thabok; Nhahin)
Acytolepis puspa gisca ; Onodera,2009b : 35,37,42. (Vientiane: Namsong Riverside; Houaphan: Phu Phan; Bolikhamxai: Phon Si)
Acytolepis puspa gisca ; Nakamura & Wakahara,2012 : 51.
Acytolepis puspa gisca ; Onodera,2022 : 75, pl.207, figs.♂,♂(Un),♀,♀(Un),♀,♀(Un). (Bolikhamxai: Tabok; Xiengkhouang: Phou Samsoum; Luang Phabang: Phou Yang)

[ CAMBODIA ]
Acytopepis puspa gisca ; Woodfield & Murton,2006 : figs.♂,♂(Un). (Botum Sakor NP.)
Acytolepis puspa ; Onodera,2009 : 40. (Siem Reap: Phnum Kulen)
Acytolepis puspa gisca ; Monastyrskii et al.,2011 : 130. (Phnom Samkos W.S.)
Acytolepis puspa gisca ; Kosterin,2019b : 29. (Ratanakiri; Mondulkiri)
Acytolepis puspa gisca ; Kosterin,2020 : 321. (Siem Reap)
Acytolepis puspa gisca ; Chartier & Kosterin,2022 : 120. (Koh Kong; Mondulkiri; Pursat; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Lycaenopsis puspa imperatrix ; Fruhstorfer,1915-1924 : 869. (Tonkin)
Cyaniris puspa imperatrix ; Godfrey,1919 : 469. (S: Daban)
Lycaenopsis puspa ; Lemée,1950 : 65. (Saigon)
Lycaenopsis puspa imperatrix ; Metaye,1957 : 91. (N / C)
Acytolepis puspa gisca ; Ikeda et al.,2001 : 48, figs.2:6,♂,7,♂(Un),8,♀,6:11(♂genitalia). (Cuc Phuong)
Acytolepis puspa gisca ; Luong et al.,2004 : 9, 75, figs.♂,♂(Un). (Cuc Phuong)
Acytolepis puspa ; Monastyrskii et al.,2006 : 42. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien)
Acytolepis puspa gisca ; Miyazaki, Saito & Saito,2006d : 53, figs.L-33,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Madagui; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Acytolepis puspa gisca ; Nihira,2009 : 101. (Phu Quoc)
Acytolepis puspa gisca ; Monastyrskii & Devyatkin,2015 : 50. (N / C / S)
Acytolepis puspa gisca ; Saito et al.,2017 : 80. (S: Nui Cam)

[ THAILAND ]
Mae Hong Son : 9♂.

Jan

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Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 14♂ 5♀ (incl. photo).

Jan

Feb

Mar

Apr

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Jul

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Oct

Nov

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Lampang : 3♂ (photo).

Jan

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Sep

Oct

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Phrae : 1♂.

Jan

Feb

Mar

Apr

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Jul

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Sep

Oct

Nov

Dec

Nan : 10♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

Oct

Nov

Dec

Uttaradit : 1♂ (photo).

Jan

Feb

Mar

Apr

May

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Oct

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Loei : 1♂ (photo).

Jan

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Apr

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Sep

Oct

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Tak : 1♂.

Jan

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Mar

Apr

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Chaiyaphum : 1♂ (photo).

Jan

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Apr

May

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Jul

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Sep

Oct

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Kanchanaburi : 1♂.

Jan

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Chonburi : 2♂.

Jan

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Chanthaburi : 12♂.

Jan

Feb

Mar

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May

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Trat : 1♂ (photo by E.Ikari).

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Phetchaburi : 2♂.

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The butterfly is very common in montane forests at low to high elevations (Alt. 30 - 1800m). Males frequently come to the banks of streams and puddles to suck water.
Some specimens of the extreme dry season form are quite similar to that of Celatoxia marginata. I consider the figured specimen of Celatoxia marginata splendens in Kimura et al. (2014) to be this taxon.

I have observed Neopithecops zalmora ♂ courting Acytolepis puspa ♂. The Neopithecops zalmora ♂ persistently courted the Acytolepis puspa ♂ and, incredibly, tried to mate with it. Eventually Acytolepis puspa ♂ flew away.

Fruhstorfer, H.,1910 : Neue Cyaniris-Rassen und Übersicht der bekannten Arten.
Stett. ent. Ztg. 71(2):282-305. (15.Mar.1910)

Fruhstorfer, H.,1917 : Revision der Lycaenidengattung Lycaenopsis auf Grund morphologischer Vergleiche der Klammerorgane.
Arch. Naturgesch. (A)82(1):1-42, 2 pls. (Jan.1917)

Corbet, A.S.,1941 : A List of the Butterflies of Hong Kong (3).
Hongkong Nat. 10(3/4):148-165A, 166, 5 figs, 1 pl.

♂, Chiang Mai, Thailand.	♀ (dsf), Chiang Mai, Thailand.
♂ (center), Chiang Mai, Thailand.	♂, Lampang, Thailand.
♂ (dsf), Chiang Mai, Thailand.

Interspecies courtship behavior, Neopithecops zalmora ♂ and Acytolepis puspa ♂
Neopithecops zalmora ♂ is courting ♂ of Acytolepis puspa, Nan, Thailand.
Neopithecops zalmora ♂ is courting ♂ of Acytolepis puspa, Nan, Thailand.

1999.06.25 - 2024.06.30.