♂, Vientiane, Laos. (photo by K.Saito)		♀, Vientiane, Laos. (photo by K.Saito)
♂, Vientiane, Laos. (photo by K.Saito)		♀, Vientiane, Laos.

ON : Euthalia apicalis lioneli Fruhstorfer, 1905 OD : Ent. Zeit. 19:(11):56. TL : Than Moi ; Chiem Hoa, "Tonkin" [N. Vietnam]. Distribution : Laos, Cambodia, Vietnam, Yunnan. ● ssp.chula ● ssp.lioneli

[ LAOS ]
Euthalia eriphylae f. lioneli ; Dubois & Vitalis,1924 : 43. (Luang-Prabang)
Euthalia eriphylae eriphylae ; Motono & Negishi,1989 : 60, pl.68, figs.5,♂,6,♂(Un),7,♀,8,♀(Un). (Vang Vieng; Tha Ngone; Vientiane)
Euthalia eriphylae lioneli ; Osada, Uémura & Uehara,1999 : 207, pl.64, figs.♂,♀. (Vang Vieng)
Euthalia eriphylae lioneli ; Nimura & Wakahara,2005 : 43. (Namdhua)
Euthalia eriphylae lioneli ; Nakamura & Wakahara,2012 : 46.
Euthalia eriphylae lioneli ; Onodera,2022 : 43, pl.127, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Khammouan: Natung; Bolikhamxai: Phon Si)

[ CAMBODIA ]
Euthalia eriphylae ; Onodera,2007 : 29, fig.70,♂. (Siem Reap: Angkor)
Euthalia eriphylae lioneli ; Chartier & Kosterin,2022 : 118. (Siem Reap)

[ VIETNAM ]
Euthalia eriphylae lioneli ; Fruhstorfer,1913 : 670. (Tonkin: Than Moi ; Chiem Hoa)
Euthalia lioneli ; Vitalis,1919 : 231. (Tonkin)
Euthalia eriphylae f. lioneli ; Dubois & Vitalis,1924 : 43. (Tonkin: Than-Moi; Chiem-Hoa)
Euthalia eriphylae lioneli ; Metaye,1957 : 97. (N)
Euthalia eriphylae lioneli ; Ikeda et al.,2000 : 36, figs.6:16,♂,17,♂(Un), fig.9:36(♂,genitalia). (Cuc Phuong)
Euthalia eriphylae lioneli ; Luong et al.,2004 : 7, 67, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Euthalia eriphylae ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Phong Dien)
Euthalia eriphylae alboapicala ; Miyazaki, Saito & Saito,2006b : 32, fig.N-87,♀(right). (Lam Dong: Dai Duc Me)
Euthalia eriphylae lioneli ; Miyazaki et al.,2010 : 21, fig.N-16,♀. (Lam Dong: Dambri; Dai Duc Me)
Euthalia eriphylae lioneli ; Monastyrskii & Devyatkin,2015 : 40. (N / C)
Euthalia eriphylae chula ; Monastyrskii & Devyatkin,2015 : 40. (S)
Euthalia eriphylae lioneli ; Monastyrskii,2019 : 4(2):117-118, pl.64, figs.1c♀,1d♀. (N: Ba Be; Huu Lien; Cat Ba; Cuc Phuong / C: Vu Quang; Phong Dien; Dakrong; Ngoc Linh / S: Nam Cat Tien; La Nga)
Euthalia alboapicala ; Monastyrskii,2019 : 4(2): pl.64, figs.1a♂,2b♂(Un). (in part)

[ LAOS ]
Vientiane : 2♂ 3♀.

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Bolikhamxai : 1♂ (photo by H.Onodera).

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Khammouan : 1♂ (photo by H.Onodera).

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[ CAMBODIA ]
Siem Reap : 2♂ (photo by H.Onodera).

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[ VIETNAM ]
Kon Tum : 1♀ (photo by K.Saito).

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Lam Dong : 6♂ 4♀ (photo by K.Saito).

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The butterfly is locally not rare in montane foreat at low to moderate elevations.
Populations in Laos and Vietnam are highly variable, both individually and regionally. In particular, populations in C. to S. Vietnam are quite variable and unstable. I treat these populations provisionally as this taxon due to insufficient comparisons, but they may be split into several taxa in the future.
I also suspect that these taxa may be subspecies of Euthalia mahadeva. Although I tentatively follow the classical classification, further taxonomic study is needed, including DNA analysis.

Fruhstorfer, H.,1905 : Neue Rhopaloceren aus dem Indo-Australischen Gebiet.
Ent. Zeit. 19(11):55-57. (15.Jun.1905)

2007.02.10 - 2023.02.28.