♂, Vientiane, Laos. (photo by K.Saito)		♀, Vientiane, Laos. (photo by K.Saito)
♂, Vientiane, Laos. (photo by K.Saito)		♀, Vientiane, Laos.

ON : Euthalia apicalis lioneli Fruhstorfer, 1905 OD : Ent. Zeit. 19:(11):56. TL : Than Moi ; Chiem Hoa, Tonkin. Distribution : Laos, Cambodia, Vietnam, Yunnan. ● ssp.chula ● ssp.lioneli

[ LAOS ]
Euthalia eriphylae f.lioneli ; Dubois & Vitalis,1924 : 43. (Luang-Prabang)
Euthalia eriphylae eriphylae ; Motono & Negishi,1989 : 60, pl.68, figs.5,♂,6,♂(Un),7,♀,8,♀(Un). (Vang Vieng; Tha Ngone; Vientiane)
Euthalia eriphylae lioneli ; Osada, Uémura & Uehara,1999 : 207, pl.64, figs.♂,♀. (Vang Vieng)
Euthalia eriphylae lioneli ; Nimura & Wakahara,2005 : 43. (Namdhua)
Euthalia eriphylae lioneli ; Nakamura & Wakahara,2012 : 46.

[ CAMBODIA ]
Euthalia eriphylae ; Onodera,2007 : 29, fig.70,♂. (Siem Reap: Angkor)

[ VIETNAM ]
Euthalia eriphylae lioneli ; Fruhstorfer,1913 : 670. (Tonkin: Than Moi ; Chiem Hoa)
Euthalia lioneli ; Dubois & Vitalis,1919 : 231. (Tonkin)
Euthalia eriphylae f.lioneli ; Dubois & Vitalis,1924 : 43. (Tonkin: Than-Moi; Chiem-Hoa)
Euthalia eriphylae lioneli ; Metaye,1957 : 97. (N)
Euthalia eriphylae lioneli ; Ikeda et al.,2000 : 36, figs.6:16,♂,17,♂(Un), fig.9:36(♂,genitalia). (Cuc Phuong)
Euthalia eriphylae lioneli ; Luong et al.,2004 : 7, 67, figs.♂,♂(Un),♀,♀(Un). (Cuc Phuong)
Euthalia eriphylae ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Phong Dien)
Euthalia eriphylae alboapicala ; Miyazaki, Saito & Saito,2006b : 32, fig.N-87,♀(right). (Lam Dong: Dai Duc Me)
Euthalia eriphylae lioneli ; Miyazaki et al.,2010 : 21, fig.N-16,♀. (Lam Dong: Dambri; Dai Duc Me)
Euthalia eriphylae lioneli ; Monastyrskii & Devyatkin,2015 : 40. (N / C)
Euthalia eriphylae chula ; Monastyrskii & Devyatkin,2015 : 40. (S)
Euthalia eriphylae lioneli ; Monastyrskii,2019 : 4(2):117-118, pl.64, figs.1c♀,1d♀. (N: Ba Be; Huu Lien; Cat Ba; Cuc Phuong / C: Vu Quang; Phong Dien; Dakrong; Ngoc Linh / S: Nam Cat Tien; La Nga)
Euthalia alboapicala ; Monastyrskii,2019 : 4(2): pl.64, figs.1a♂,2b♂(Un). (in part)

[ LAOS ]
Vientiane : 2♂ 3♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Bolikhamxai : 1♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Khammouan : 1♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ CAMBODIA ]
Siem Reap : 2♂ (photo by H.Onodera).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ VIETNAM ]
Kon Tum : 1♀ (photo by K.Saito).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Lam Dong : 6♂ 4♀ (photo by K.Saito).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is locally not rare in montane foreat at low to moderate elevations.
Populations in Laos and Vietnam are highly variable in individual and regional variations. In particular, populations in C. to S.Vietnam are quite variable and unstable. I treat these populations as this taxon tentatively due to insufficient comparison, however may be divided into several taxa in the future.
Furthermore I suspect these taxa might be subspecies of Euthalia mahadeva. Although I follow the classical classification provisionally, further taxonomic study is needed including DNA analysis.

Fruhstorfer, H.,1905 : Neue Rhopaloceren aus dem Indo-Australischen Gebiet.
Ent. Zeit. 19(11):55-57.

2007.02.10 - 2021.09.30.