Moduza procris procris  (Cramer,[1777])

Moduza procris procris ♂ Up. Moduza procris procris ♂ Un. Moduza procris procris ♀ Up. Moduza procris procris ♀ Un.
♂ (wsf), Chiang Mai, Thailand. ♀ (wsf), Chiang Mai, Thailand.
Moduza procris procris ♂ Up. Moduza procris procris ♂ Un. Moduza procris procris ♀ Up. Moduza procris procris ♀ Un.
♂ (dsf), Chanthaburi, Thailand. ♀ (dsf), Chiang Mai, Thailand.
Moduza procris procris ♂ Up. Moduza procris procris ♂ Up. Moduza procris procris ♀ Up. Moduza procris procris ♀ Up.
♂ (wsf), Phetchaburi, Thailand. ♀ (dsf), Champasak, Laos.
Moduza procris procris ♂ Up. Moduza procris procris ♂ Un. Moduza procris procris ♂ Up. Moduza procris procris ♂ Un.
♂, Dong Nai, Vietnam. (photo by K.Saito) ♂ (ab.), Nan, Thailand.

ON : Papilio procris Cramer,[1777]
OD : Uitl. Kapellen 2(9-16):15, pl.106, figs.E,F.
TL : China.




Distribution :
Nepal, India to Myanmar, Thailand, Laos, Cambodia, Vietnam,
S.China, Hainan.
procris map

Synonyms

Limenitis procris procris forma temp.chlaena Fruhstorfer,1906 : Ent. Zeit. 20(9):62. TL. Sikkim.
Modusa [sic] neoprocris : Chou, Yuan, Yin, Zhang & Chen,2002 : Entomotaxonomia 24(1):54, 58, figs.14a,(HT,♂),14b,(HT,♂Un). TL. Fujian, China.
Moduza procris hainana : Tsukada,1991 : Butterflies of the South East Asian Islands. 5:317,545, pl.234, figs.1(PT,♂),2(HT,♂). TL. Hainan. (♂, in coll. E.Tsukada)


Type Material Information

neoprocris : Holotype ♂, Quanzhou, Fujian. (Entomological Museum, North west Sci-Tech University)
hainana : Holotype ♂, 1988.Aug.10. Tong-za, Hainan. (in coll. E.Tsukada) ; Paratype 1♂, 1988.Feb.26. Hainan. (in coll. E.Tsukada)


Records taken from Literature

[ THAILAND ]
Limenitis procris procris ; Fruhstorfer,1913 : 640. (Siam)
Limenitis procris procris ; Godfrey,1916 : 129-130. (Petchaburi River)
Limenitis procris procris ; Godfrey,1927 : 124. (N)
Limenitis procris procris ; Godfrey,1930 : 287. (except P)
Moduza procris procris ; Pinratana,1979 : 63, figs.N111,♂,♂(Un). (Chiang Mai / Chanthaburi / Chonburi / Kao Yai / Kanchanaburi / Phitsanulok / Ratchaburi)
Moduza procris procris ; Pinratana & Eliot,1996 : 64, pl.41, fig.116a,♂. (Chiang Mai / Phisanulok / Nakhon Nayok / Chanthaburi / Chonburi / Kanchanaburi / Ratchaburi)
Moduza procris procris ; Yamaguchi & Aoki,2000 : 255, fig.101,♂. (Mukdahan: Phu Mu ; Phu Pha Thaep ; Phu Pha Phung)
Moduza procris procris ; Yamaguchi & Aoki,2002 : 28. (Mukdahan: Ban Na Lak)
Moduza procris procris ; Yamaguchi, Aoki & Akiyama,2005 : 51. (Mukdahan: Phu Huai Sing / Ubon Ratchathani: Phu Chong Na Yoi)
Moduza procris procris ; Katsuyama et al.,2005 : 70, fig.163,♂. (Umphang)
Moduza procris procris ; Ek-Amnuay,[2007] : 414, pl.181, figs.N121a,♂,♂(Un),♀,♂(ab.),♂(ab.Un). (Chiang Mai / Khao Yai)
Moduza procris procris ; Ek-Amnuay,2012 : 424, pl.185, figs.N287a,♂,♂(Un),♀,♂(ab.),♂(ab.Un) (Chiang Mai / Khao Yai)
Moduza procris procris ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Moduza procris procris ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Sa Kaeo / Nakhon Nayok / Chanthaburi / Trat) (accessed: 2020.09.24)
Moduza procris procris ; Kimura et al.,2016 : 93, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Chiang Dao / Doi Suthep / Ban Mae Wa / Thung Saleng Luang / Phu Kradung / Phu Khiewo / Kao Yai / Sarika / Nakohn Nayok / Wan Nam Khiewo / Kao Soi Dao / Kratchin WF / Pliewn WF / Kao Kiewo / Sai York Noi)

[ LAOS ]
Moduza procris procris ; Motono & Negishi,1989 : 59, pl.63, figs.5,♂,6,♂(Un),7,♀,8,♀(Un). (Vang Vieng / Nam Moh / Nam Ngum Dam)
Moduza procris procris ; Osada, Uémura & Uehara,1999 : 207, pl.61, figs.♂,♀. (Thabok / Xaignabouri)
Moduza procris procris ; Nimura & Wakahara,2005 : 42. (Namdhua)
Moduza procris procris ; Onodera,2009b : 35,41. (Vientiane: Namsong Riverside / Khammouan: Nacay)
Moduza procris procris ; Nakamura & Wakahara,2012 : 46.

[ CAMBODIA ]
Moduza procris procris ; Woodfield & Murton,2006 : figs,♂,♂(Un). (Botum Sakor NP.)
Moduza procris ; Onodera,2009 : 64. (Kratie: Snoul)

[ VIETNAM ]
Limenitis procris procris ; Fruhstorfer,1913 : 640. (Tonkin)
Limenitis procris ; Lemee,1950 : 14. (Hagiang / Hanoi)
Limenitis procris procris ; Metaye,1957 : 97. (N / C / S)
Moduza procris procris ; Ikeda et al.,2000 : 36, figs.6:6,♂,7,♂(Un), fig.9:34(♂,genitalia). (Cuc Phuong)
Moduza procris ; Lien,2003 : 57. (Phu Quoc)
Moduza procris procris ; Hao et al.,2004 : 7, 65, figs.♂[♀],♂[♀](Un),♀[♂]. (Cuc Phuong)
Moduza procris ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong; A Luoi; Phong Dien; Bach Ma)
Moduza procris procris ; Miyazaki, Saito & Saito,2006b : 26, figs.N-76,♂,♀. (Lam Dong: Dambri ; Dai Duc Me / Dong Nai: Thac Mai)
Moduza procris procris ; Monastyrskii & Devyatkin,2015 : 36. (N / C / S)
Moduza procris procris ; Saito et al.,2017 ; 81. (S: Nui Coto)
Moduza pricris pricris ; Monastyrskii,2019 : 2:43-44, pl.39, figs.3a,♂,3b,♂(Un). (N: Muong Nhe ; Na Hang ; Ba Be ; Tam Dao ; Cuc Phuong / C: Ben En ; Pu Hoat ; Pu Mat ; Huong Son ; Vu Quang ; Phong Dien ; Dakrong ; Bach Ma ; Ba Na ; Son Thanh ; Ba To ; Kon Cha Rang ; Bu Gia Map ; Loc Bao ; Nui Chua ; Da Teh ; La Nga ; Bao Lam / S: Cat Tien ; Con Dao)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 5♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 6♂ 6♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 6♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Nakhon Nayok : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chonburi : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 4♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Champasak : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Lam Dong : 3♂ 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Dong Nai : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
The butterfly is common in low to moderate elevations (alt.200m - 1200m).
Tsukada (1991) described ssp.hainana from Hainan mainly due to the large cell white spot of the UpF. However, the size of this spot is very variable and the individuals have this characteristics are occasionally found in continental China to Indochina region especially during in the dry season. Therefore, I consider it should be included in nominotypical subspecies.

Literature

Cramer, P.,[1777] :
Uitlandsche Kapellen. voorkomende in de drie Waereld-Deelen Asia, Africa en America, by een verzameld en bescreeven.
Amsterdam. Baalde, and Utrecht, Wild. 2(9-16):1-152, pl.97-192.

Fruhstorfer, H.,1906 : Neue und seltene indo-malayische Rhopaloceren.
Ent. Zeit. 20(9):61-63.

Tsukada, E.,1991 : Butterflies of the South East Asian Islands V, Nymphalidae (II). (in Japanese)
Azumino Butterflie's Research Institute. 576pp.(238 col. pls.)

Chou, Yuan, Yin, Zhang & Chen,2002 : New Species, New Subspecies, and New Records of Butterflies from China (VI).
Entomotaxonomia 24(1):52-68, 23 figs.

Lang, S.,2009 : A Study on the systematics of Nymphalidae (excepting Satyrinae) from China (Lepidoptera: Papilionoidea)
Beijing, P.R.China.

Moduza procris procris ♂ Moduza procris procris ♂
♂ (wsf), Chiang Mai, Thailand. ♂ (dsf), Chiang Mai, Thailand.
Moduza procris procris ♂ Moduza procris procris ♂
♂ (dsf), Chiang Mai, Thailand. ♂, Chiang Mai, Thailand.

2007.03.10 - 2021.02.28.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © Y.Inayoshi.