♂ (dsf), Chiang Mai, Thailand.		♀ (dsf), Mae Hong Son, Thailand.
♂ (dsf), Tak, Thailand.		♀ (dsf), Trat, Thailand.
♂, Chanthaburi, Thailand.		♀, Mae Hong Son, Thailand.
♂, Ha Giang, Vietnam. (photo by K.Saito)		♀ (dsf), Chiang Mai, Thailand.

ON : Neptis kamarupa Moore, 1875 OD : Proc. zool. Soc. Lond. 1874(4):570. TL : Assam, [India]. (Oxford.) Distribution : India (Assam) to Myanmar, Thailand, Laos, Cambodia, Vietnam, SE.Tibet, S.Yunnan, SW.Guangxi. ● ssp.kamarupa ● ssp.papaja

Neptis adara Moore, 1879 : Proc. zool. Soc. Lond. 1878(4):830. TL. Moulmein to Meetan ; Ahsoun ; Naththoung to Paboga ; Moolai, [Myanmar]. (NHML)
Neptis meetana Moore, 1879 : Proc. zool. Soc. Lond. 1878(4):830. TL. Meetan ; Taoo, [Myanmar]. (NHML)

[ THAILAND ]
Neptis hylas acerides ; Godfrey,1916 : 128.
Neptis hylas hylas ; Godfrey,1927 : 123. (N)
Neptis hylas hylas ; Godfrey,1930 : 279. (except P)
Neptis hylas acerides ; Davidson & Macbeth,1938 : 74. (N: Koon Tarn)
Neptis hylas kamarupa ; Eliot,1969 : 61. (Siam)
Neptis hylas kamarupa ; Pinratana,1979 : 45, figs.N72a♂,♂(Un). (Chiang Mai; Chaiyaphum; Chanthaburi; Kanchanaburi; Nakhon Nayok; Nonthaburi; Saraburi)
Neptis hylas kamarupa ; Pinratana & Eliot,1996 : 47, pl.27, figs.65b♂,♂(Un). (Chiang Mai; Loei; Chaiyaphum; Nakhon Ratchasima; Nakhon Nayok; Saraburi; Chanthaburi; Kanchanaburi)
Neptis hylas kamarupa ; Yamaguchi & Aoki,2000 : 255, fig.95,♂. (Mukdahan: Phu Mu; Phu Pha Thaep; Phu Huay Sing)
Neptis hylas kamarupa ; Yamaguchi & Aoki,2002 : 27. (Ubon Ratchathani: Ban Ba Hai; Ban Na Pho Krang; Phu Chong Na Yoi; Nam Tok Soi Sawan; Kaeng E-Keow; Mukdahan: Ban Lao Kram; Phu Huay Sing; Nam Tok Tad Ton; Ban Na Lak; Ban Gok Tum)
Neptis hylas kamarupa ; Yamaguchi, Aoki & Akiyama,2005 : 50. (Mukdahan: Ban Lao Kram; Ban Na Lak; Phu Hin Khan; Phu Huai Sing; Nam Tok Tad Ton; Ubon Ratchathani: Warin Chamrap; Nam Tok Soi Sawan; Phu Chong Na Yoi; Ban Ba Hai; Phu Pung; Don Far Huan)
Neptis hylas kamarupa ; Katsuyama et al.,2005 : 69, fig.151,♂. (Umphang: Umphang; Tha Sai)
Neptis hylas kamarupa ; Katsuyama et al.,2006 : 49, fig.113,♂. (Nan: Mae Charim; Ban Nam Poon)
Neptis hylas kamarupa ; Ek-Amnuay,[2007] : 388, pl.168, figs.N68b♀,♀(Un). (in part)
Neptis hylas kamarupa ; Ek-Amnuay,2012 : 398,400, pl.178, figs.N243a♀,♀(Un). (in part)
Neptis hylas kamarupa ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Neptis hylas kamarupa ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs., Early Stages. (Chiang Mai; Tak; Kanchanaburi; Sa Kaeo; Chanthaburi; Ratchaburi) (accessed: 2023.06.13)
Neptis hylas kamarupa ; Kimura et al.,2016 : 74-75, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Doi Ankhaan; Chiang Dao; Doi Phui; Doi Suthep; Doi Inthanon; Doi Pa Chang; Thung Saleng Luang; Phu Kradung; Phu Khiewo; Kao Yai; Sarika; Nakohn Nayok; Wan Nam Khiewo; Kao Soi Dao; Kratching WF; Kao Chamao; Kao Khiewo; Bang Phra; Wan Khiewo; Tha Sai; Sai York Noi; Erawan WF; Kanchanaburi; Ratchaburi; Kaeng Krachang)
Neptis hylas kamarupa ; Chaianunporn & Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Neptis hylas f. acerides ; Dubois & Vitalis,1924 : 39. (Laos)
Neptis hylas kamarupa ; Motono & Negishi,1989 : 57, pl.56, figs.1,♂,2,♂(Un),3,♀,4,♀(Un). (Luang Prabang; Tha Ngone; Nam Moh; Vientiane; Ban Houei Sai; Nam Ngum Dam)
Neptis hylas kamarupa ; Osada, Uémura & Uehara,1999 : 206, pl.54, figs.♂,♂(Un),♀. (Xam Neua; Vieng Xai)
Neptis hylas kamarupa ; Nimura & Wakahara,2005 : 42. (Nakai)
Neptis hylas kamarupa ; Onodera,2009b : 35. (Vientiane: Namsong Riverside)
Neptis hylas kamarupa ; Nakamura & Wakahara,2012 : 45.
Neptis hylas kamarupa ; Onodera,2022 : 33, pl.108, figs.♂,♂(Un),♀,♀(Un). (Oudomxai; Xiengkhouang: Tham Piou)

[ CAMBODIA ]
Neptis hylas ; Onodera,2007 : 22, fig.53,♂. (Rattanakiri: Ta Veaeng)
Neptis hylas kamarupa ; Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)
Neptis hylas kamarupa ; Kosterin,2019 : 97. (Kep)
Neptis hylas kamarupa ; Kosterin,2019b : 32. (Ratanakiri; 'Mondulkiri')
Neptis hylas kamarupa ; Kosterin,2020 : 328. (Siem Reap; Preah Vihear; Stung Treng)
Neptis hylas kamarupa ; Chartier & Kosterin,2022 : 117. (Battambang; Kep; Koh Kong; Kampong Thom; Mondulkiri; Phnom Penh; Preah Sihanouk; Pursat; Preah Vihear; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Neptis adara ; Joannis,1901 : 320. (Tonkin: Cao Bang)
Neptis hylas acerides ; Godfrey,1919 : 467. (S: Daban)
Neptis acerides ; Vitalis,1919 : 229. (Annam / Tonkin)
Neptis hylas f. luculenta ; Dubois & Vitalis,1924 : 39. (Tonkin)
Neptis hylas f. acerides ; Dubois & Vitalis,1924 : 39. (Tonkin / Annam)
Neptis hylas ; Lemée,1950 : 14. (Hagiang)
Neptis hylas hylas ; Metaye,1957 : 96. (N / C / S)
Neptis hylas luculenta ; Metaye,1957 : 96. (N)
Neptis hylas acerides ; Metaye,1957 : 96. (N / C)
Neptis hylas kamarupa ; Eliot,1969 : 61. (N / S)
Neptis hylas hylas ; Ikeda et al.,2000 : 36, fig.5:3,♂,4,♂(Un),5,♀, fig.9:27(♂,genitalia). (Cuc Phuong)
Neptis hylas ; Vu,2003 : 57. (Phu Quoc)
Neptis hylas kamarupa ; Luong et al.,2004 : 6, 62, figs.♂[♀],♂[♀](Un). (Cuc Phuong)
Neptis hylas ; Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Bach Ma)
Neptis hylas ; Miyazaki, Saito & Saito,2006b : 20, figs.N-44,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Neptis kamarupa ; Miyazaki, Saito & Saito,2006b : 20, figs.N-45,♂,♀. (Lam Dong: Dambri; Gung Re)
Neptis hylas kamarupa ; Nihira,2009 : 99. (Phu Quoc)
Neptis hylas kamarupa ; Yokochi et al.,2015 : 40. (Khanh Hoa: Hon Ba)
Neptis hylas kamarupa ; Monastyrskii & Devyatkin,2015 : 37. (N / C / S)
Neptis hylas kamarupa ; Saito et al.,2017 : 81. (S: Nui Cam)
Neptis hylas kamarupa ; Monastyrskii,2019 : 4(2):62-63, pl.43, figs.2a♂,2b♂(Un),2c♂,2d♂(Un). (N / C / S)

[ THAILAND ]
Mae Hong Son : 3♂ 6♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 15♂ 2♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 3♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Loei : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchabun : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chaiyaphum : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chonburi : 4♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Trat : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Bolikhamxai : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Vientiane : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ VIETNAM ]
Ha Giang : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is very common in montane and secondary forests at low to high elevations (Alt. 10 - 1600m) and is often found on forest edges and forest paths.
The populations from indo-china region are very variable as Kimura et al. (2016) pointed out. Further taxonomic study is needed.

Moore, F.,1875 : Descriptions of New Asiatic Lepidoptea.
Proc. zool. Soc. Lond. 1874(4):565-579, pls.66,67.

Moore, F.,1879 : A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim, with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4):821-858, pls.51-53.

Eliot, J.N.,1969 : An Analysis of the Eurasian and Australian Neptini (Lepidoptera: Nymphalidae).
Bull. Br. Mus. nat. Hist. (Ent.) Supple.15. London.

♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.

2006.02.20 - 2024.02.29.