♂ (dsf), Chiang Mai, Thailand.		♀ (dsf), Mae Hong Son, Thailand.
♂ (dsf), Tak, Thailand.		♀ (dsf), Trat, Thailand.
♂, Chanthaburi, Thailand.		♀, Mae Hong Son, Thailand.
♂, Ha Giang, Vietnam. (photo by K.Saito)		♀ (dsf), Chiang Mai, Thailand.

ON : Neptis kamarupa Moore, 1875 OD : Proc. zool. Soc. Lond. 1874(4): 570. TL : Assam. (Oxford.) Distribution : Assam to Myanmar, S.E.Tibet, S.Yunnan, Thailand, Laos, Cambodia, Vietnam, S.W.Guangxi. ● ssp.kamarupa ● ssp.papaja

Neptis adara Moore, 1879; Proc. zool. Soc. Lond. 1878(4): 830; TL: 'Moulmein to Meetan; Ahsoun; Naththoung to Paboga; Moolai, [Upper Tenasserim]' [Kayin State, Myanmar]. (NHML)
Neptis meetana Moore, 1879; Proc. zool. Soc. Lond. 1878(4): 830; TL: 'Meetan; Taoo, [Upper Tenasserim]' [Kayin State, Myanmar]. (NHML)

[ THAILAND ]
Neptis hylas acerides: Godfrey,1916 : 128.
Neptis hylas hylas: Godfrey,1927 : 123. (N)
Neptis hylas hylas: Godfrey,1930 : 279. (except P)
Neptis hylas acerides: Davidson & Macbeth,1938 : 74. (N: Koon Tarn)
Neptis hylas kamarupa: Eliot,1969 : 61. (Siam)
Neptis hylas kamarupa: Pinratana,1979 : 45, figs.N72a♂,♂(Un). (Chiang Mai; Chaiyaphum; Chanthaburi; Kanchanaburi; Nakhon Nayok; Nonthaburi; Saraburi)
Neptis hylas kamarupa: Pinratana & Eliot,1996 : 47, pl.27, figs.65b♂,♂(Un). (Chiang Mai; Loei; Chaiyaphum; Nakhon Ratchasima; Nakhon Nayok; Saraburi; Chanthaburi; Kanchanaburi)
Neptis hylas kamarupa: Yamaguchi & Aoki,2000 : 255, fig.95,♂. (Mukdahan: Phu Mu; Phu Pha Thaep; Phu Huay Sing)
Neptis hylas kamarupa: Yamaguchi & Aoki,2002 : 27. (Ubon Ratchathani: Ban Ba Hai; Ban Na Pho Krang; Phu Chong Na Yoi; Nam Tok Soi Sawan; Kaeng E-Keow; Mukdahan: Ban Lao Kram; Phu Huay Sing; Nam Tok Tad Ton; Ban Na Lak; Ban Gok Tum)
Neptis hylas kamarupa: Yamaguchi, Aoki & Akiyama,2005 : 50. (Mukdahan: Ban Lao Kram; Ban Na Lak; Phu Hin Khan; Phu Huai Sing; Nam Tok Tad Ton; Ubon Ratchathani: Warin Chamrap; Nam Tok Soi Sawan; Phu Chong Na Yoi; Ban Ba Hai; Phu Pung; Don Far Huan)
Neptis hylas kamarupa: Katsuyama et al.,2005 : 69, fig.151,♂. (Umphang: Umphang; Tha Sai)
Neptis hylas kamarupa: Katsuyama et al.,2006 : 49, fig.113,♂. (Nan: Mae Charim; Ban Nam Poon)
Neptis hylas kamarupa: Ek-Amnuay,[2007] : 388, pl.168, figs.N68b♀,♀(Un). (partim)
Neptis hylas kamarupa: Ek-Amnuay,2012 : 398,400, pl.178, figs.N243a♀,♀(Un). (partim)
Neptis hylas kamarupa: Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Neptis hylas kamarupa: Sophonviwatkul et al.,2015– : Butterflies of Thailand, figs., Early Stages. (Chiang Mai; Tak; Kanchanaburi; Sa Kaeo; Chanthaburi; Ratchaburi) (accessed: 2023.06.13)
Neptis hylas kamarupa: Kimura et al.,2016 : 74–75, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un),♀,♀(Un). (Doi Ankhaan; Chiang Dao; Doi Phui; Doi Suthep; Doi Inthanon; Doi Pa Chang; Thung Saleng Luang; Phu Kradung; Phu Khiewo; Kao Yai; Sarika; Nakohn Nayok; Wan Nam Khiewo; Kao Soi Dao; Kratching WF; Kao Chamao; Kao Khiewo; Bang Phra; Wan Khiewo; Tha Sai; Sai York Noi; Erawan WF; Kanchanaburi; Ratchaburi; Kaeng Krachang)
Neptis hylas kamarupa: K. Chaianunporn & T. Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)
Neptis hylas: Chueamueangphan et al.,2025 : 208. (Phayao: Mae Puem N.P.)

[ LAOS ]
Neptis hylas f. acerides: Dubois & Vitalis,1924 : 39. (Laos)
Neptis hylas kamarupa: Motono & Negishi,1989 : 57, pl.56, figs.1,♂,2,♂(Un),3,♀,4,♀(Un). (Luang Prabang; Tha Ngone; Nam Moh; Vientiane; Ban Houei Sai; Nam Ngum Dam)
Neptis hylas kamarupa: Osada et al.,1999 : 206, pl.54, figs.♂,♂(Un),♀. (Xam Neua; Vieng Xai)
Neptis hylas kamarupa: Nimura & Wakahara,2005 : 42. (Nakai)
Neptis hylas kamarupa: Onodera,2009b : 35. (Vientiane: Namsong Riverside)
Neptis hylas kamarupa: Nakamura & Wakahara,2012 : 45.
Neptis hylas kamarupa: Onodera,2022 : 33, pl.108, figs.♂,♂(Un),♀,♀(Un). (Oudomxai; Xiengkhouang: Tham Piou)

[ CAMBODIA ]
Neptis hylas: Onodera,2007 : 22, fig.53,♂. (Rattanakiri: Ta Veaeng)
Neptis hylas kamarupa: Monastyrskii et al.,2011 : 129. (Phnom Samkos W.S.)
Neptis hylas kamarupa: Kosterin,2019 : 97. (Kep)
Neptis hylas kamarupa: Kosterin,2019b : 32. (Ratanakiri; 'Mondulkiri')
Neptis hylas kamarupa: Kosterin,2020 : 328. (Siem Reap; Preah Vihear; Stung Treng)
Neptis hylas kamarupa: Chartier & Kosterin,2022 : 117. (Battambang; Kep; Koh Kong; Kampong Thom; Mondulkiri; Phnom Penh; Preah Sihanouk; Pursat; Preah Vihear; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Neptis adara: Joannis,1901 : 320. (Tonkin: Cao Bang)
Neptis hylas acerides: Godfrey,1919 : 467. (S: Daban)
Neptis acerides: Vitalis,1919 : 229. (Annam; Tonkin)
Neptis hylas f. luculenta: Dubois & Vitalis,1924 : 39. (Tonkin)
Neptis hylas f. acerides: Dubois & Vitalis,1924 : 39. (Tonkin; Annam)
Neptis hylas: Lemée,1950 : 14. (Hagiang)
Neptis hylas hylas: Metaye,1957 : 96. (N; C; S)
Neptis hylas luculenta: Metaye,1957 : 96. (N)
Neptis hylas acerides: Metaye,1957 : 96. (N; C)
Neptis hylas kamarupa: Eliot,1969 : 61. (N; S)
Neptis hylas hylas: Ikeda et al.,2000 : 36, fig.5:3,♂,4,♂(Un),5,♀, fig.9:27(♂,genitalia). (Cuc Phuong)
Neptis hylas: Vu,2003 : 57. (Phu Quoc)
Neptis hylas kamarupa: Luong et al.,2004 : 6, 62, figs.♂[♀],♂[♀](Un). (Cuc Phuong)
Neptis hylas: Monastyrskii et al.,2006 : 41. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Bach Ma)
Neptis hylas: Miyazaki et al.,2006b : 20, figs.N-44,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Neptis kamarupa: Miyazaki et al.,2006b : 20, figs.N-45,♂,♀. (Lam Dong: Dambri; Gung Re)
Neptis hylas kamarupa: Nihira,2009 : 99. (Phu Quoc)
Neptis hylas kamarupa: Yokochi et al.,2015 : 40. (Khanh Hoa: Hon Ba)
Neptis hylas kamarupa: Monastyrskii & Devyatkin,2015 : 37. (N; C; S)
Neptis hylas kamarupa: Saito et al.,2017 : 81. (S: Nui Cam)
Neptis hylas kamarupa: Monastyrskii,2019 : 4(2):62–63, pl.43, figs.2a♂,2b♂(Un),2c♂,2d♂(Un). (N; C; S)
Neptis hylas kamarupa: Saito & Vu,2020a : 177. (Ha Giang: Dong Van)
Neptis hylas kamarupa: Saito et al.,2021 : 464. (Phu Quoc)

[ THAILAND ]
Mae Hong Son : 3♂ 6♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chiang Mai : 15♂ 2♀ (incl. photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Nan : 3♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Loei : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Tak : 3♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Phetchabun : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chaiyaphum : 1♂ (photo).

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chonburi : 4♂ 2♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Chanthaburi : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Trat : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ LAOS ]
Bolikhamxai : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

Vientiane : 1♂.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

[ VIETNAM ]
Ha Giang : 2♂ 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

Aug

Sep

Oct

Nov

Dec

The butterfly is very common in forests at low to high elevations (10–1,600 m a.s.l.) and is often found on forest edges and forest paths.
The populations from indo-china region are very variable as Kimura et al. (2016) pointed out. Further taxonomic study is needed.

Moore, F, 1875. Descriptions of New Asiatic Lepidoptea.
Proceedings of the zoological Society of London, 1874(4): 565–579, pls. 66,67.

Moore, F, 1879. A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim, with descriptions of new species.
Proceedings of the zoological Society of London, 1878(4): 821–858, pls. 51–53.

Eliot, J.N, 1969. An Analysis of the Eurasian and Australian Neptini (Lepidoptera: Nymphalidae).
Bulletin of the British Museum (Natural History), Entomology Supplement 15: 1–155, pls. 1–3.

♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.

2006.02.20 - 2026.03.31.