♂, Chiang Mai, Thailand.		♀, Nan, Thailand.
♂, Ranong, Thailand.		♀, Satun, Thailand.
♂, Chanthaburi, Thailand.		♂, Ranong, Thailand.

ON : Tagiades maura Snellen, 1880 OD : Midden-Sumatra 4(1)(8): 28-29. TL : Sumatra. Distribution : S.Myanmar, Thailand, Laos, Cambodia, S.Vietnam, Langkawi, W.Malaysia, Tioman, Singapore, Sumatra, Sipora, Bangka, Nias. ● ssp.maura ● ssp.diocles

Kerana diocles evaira Fruhstorfer, 1911 : Duet. ent. Zeit. [Iris] 25(2): 10. TL. "Siam" [Thailand]. (NHML)
Kerana diocles sumata Fruhstorfer, 1911 : Deut. ent. Zeit. [Iris] 25(2): 10. TL. Insel Nias. (NHML)

[ THAILAND ]
Kerana diocles evaira ; Godfrey,1916 : 145.
Tamela diocles ; Pendlebury,1923 : 48. (Nakhon Sri Thammarat: Tai Sai River; Khao Ram)
Kerana nigrita ; Godfrey,1930 : 371. (Pak Jong; Chonburi: Sriracha; Me Wong)
Kerana nigrita ; Godfrey,1932a : 275. (Trang: Chawng)
Ancistroides nigrita maura ; Evans,1949 : 280. (Siam)
Ancistroides nigrita maura ; Shirôzu & Saigusa,1962 : 34-35, pl.2, [fig.10,♂ as Iambrix salsara], Textfig.7,♂genitalia. (Mae Wang)
Ancistroides nigrita maura ; Kawazoé,1964 : 128-129. (Khao Chong)
Ancistroides nigrita diocles ; Pinratana,1985 : 63, pl.21, fig.125a♂. (Chiang Mai; Nakhon Nayok)
Ancistroides nigrita maura ; Pinratana,1985 : 63, pl.21, fig.125b♂. (Chanthaburi; Trang; Yala)
Ancistroides nigrita maura ; Yamaguchi & Aoki,2000 : 250, fig.17,♂. (Mukdahan: Ban Naudom)
Ancistroides nigrita maura ; Yamaguchi & Aoki,2002 : 12. (Ubon Ratchathani: Phu Chong Na Yoi; Mukdahan: Phu Huay Sing; Ban Lao Kram; Ban Na Lak; Ban Gok Tum)
Stimula swinhoei swinhoei ; Ek-Amnuay,[2007] : Pl.360, fig.H150,♀(Un). (in part)
Ancistroides nigrita diodes [sic] ; Ek-Amnuay,[2007] : 786, Pl.360, fig.H151a♀. (Chiang Mai)
Ancistroides nigrita othonias ; Ek-Amnuay,[2007] : 786, pl.360, figs.H151b♂(Un),♀[♂],♀[♂](Un). (Yala)
Ancistroides nigrita othonias ; Les Day,2011-2022 : Samui Butterflies, figs.Up,Un. (Koh Samui) (accessed: 2022.02.11)
Ancistroides nigrita diocles ; Kimura et al.,2011 : 65, figs.♂,♂(Un). (Doi Par Hom Pok; Doi Suthep; Doi Inthanon)
Ancistroides nigrita maura ; Kimura et al.,2011 : 65, figs.♂,♂(Un),♀,♀(Un). (Phu Khiewo; Kao Yai; Nakohn Nayok; Kao Khiewo; Kao Soi Dao; Kratching WF; Pliewn WF; Kao Chamaoh; Kam Puan; Khlong Naka; Ka Chong; Satun; Than To WF.)
Stimura swinhoei swinhoei ; Ek-Amnuay,2012 : pl.387, fig.H146,♀(Un). (in part).
Ancistroides nigrita diocles ; Ek-Amnuay,2012 : 838, pl.387, figs.H147a♂,♂(Un). (Chiang Mai)
Ancistroides nigrita othonias ; Ek-Amnuay,2012 : 838, pl.387, figs.H147b♂(Un),♀,♀(Un). (Yala)
Ancistroides nigrita maura ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Ancistroides nigrita maura ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Sa Kaeo; Nakhon Nayok; Chanthaburi) (accessed: 2023.08.04)
Stimura swinhoei ; Küppers & Janikorn,2017 : 95, figs.Abb.H-141-142. (Phang Nga)
Ancistroides nigrita othonias ; Küppers & Janikorn,2017 : 95-96, figs.Abb.H-143-146,304. (Phang Nga)

[ LAOS ]
Ancistroides nigrita maura ; Motono & Negishi,1989 : 82, pl.114, figs.B3,♂,B4,♂(Un). (Tha Ngone; Luang Prabang; Tha Dua)
Ancistroides nigrita maura ; Osada, Uémura & Uehara,1999 : 223, pl.139, fig.♂. (Nhahin)
Ancistroides nigrita diocles ; Nakamura & Wakahara,2012 : 59.
Ancistroides nigrita diocles ; Onodera,2022 : 115, pl.251, figs.♂,♂(Un),♀,♀(Un). (Khammouan: Nahin; Savannakhet)

[ CAMBODIA ]
Ancistroides nigrita maura ; Kosterin,2019 : 99, figs.59(Un),60(Un),61(Un). (Koh Kong; Preah Sihanouk)
Ancistroides nigrita maura ; Kosterin,2019b : 34. (Mondulkiri)
Ancistroides nigrita maura ; Kosterin,2020 : 332. (Siem Reap)
Ancistroides nigrita maura ; Chartier & Kosterin,2022 : 125. (Koh Kong; Mondulkiri; Preah Sihanouk; Siem Reap)

[ VIETNAM ]
Ancistroides nigrita maura ; Inoué & Kawazoé,1966 ; 98-99, fig.90,♂. (Phu Lam; Trang Bom; Djiring; Col de Blao; Banmethuot)
Ancistroides nigrita maura ; Miyazaki, Saito & Saito,2007b : 11, figs.H-67,♂,♀. (Lam Dong: Dambri; Dai Duc Me)
Ancistroides nigrita maura ; Monastyrskii & Devyatkin,2015 : 79. (S)
Ancistroides nigrita maura ; Saito et al.,2017 : 76. (S: Nui Cam; Nui Coto)

[ THAILAND ]
Mae Hong Son : 4♂ 1♀.

Jan

Feb

Mar

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May

Jun

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Aug

Sep

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Nov

Dec

Chiang Mai : 6♂ (incl. photo).

Jan

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Mar

Apr

May

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Sep

Oct

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Dec

Lampang : 1♂.

Jan

Feb

Mar

Apr

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Sep

Oct

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Phrae : 1♂.

Jan

Feb

Mar

Apr

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Jul

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Oct

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Nan : 5♂ 1♀.

Jan

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Oct

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Tak : 1♀.

Jan

Feb

Mar

Apr

May

Jun

Jul

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Sep

Oct

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Nakhon Nayok : 1♂ 1♀ (photo).

Jan

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Sep

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Sa Kaeo : 1♂ (photo).

Jan

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Oct

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Chanthaburi : 3♂.

Jan

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Mar

Apr

May

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Sep

Oct

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Phetchaburi : 1♂.

Jan

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Ranong : 6♂.

Jan

Feb

Mar

Apr

May

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Oct

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Trang : 2♂ 1♀.

Jan

Feb

Mar

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May

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Satun (incl. Koh Tarutao) : 4♂ 1♀.

Jan

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Yala : 4♂ (incl. photo).

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The butterfly is locally common in montane forests at low to modelate elevations (Alt. 10 - 1300m). Both sexes visit flowers.
Zhang et al. (2022: 39) suggested that the taxon should be treated in genus Tamela based on molecular analysis. Subsequently, Zhang et al. (2023: 17) treated both taxa diocles and maura as distinct species based on molecular analysis. However, important populations in the Indochina region have not been examined. I treat this as conventionally until this is resolved. Further taxonomic study is needed.

Snellen, P.C.T.,1880 : Natuurlijke Historie. Achtse afdeeling. Lepidoptera door P. C. T. Snellen, met eene inleiding door Joh. F. Snelleman.
In: Veth, P.J., Midden-Sumatra. Reizen en onderzoekeningen der Sumatra-Expeditie, uitgerust door het Aardrijkskundig Genootschap,1877-1879. Leiden, E.J. Brill.
4(1)(8): 1-84 (1880).

Fruhstorfer, H.,1911 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 25(2): 9-24. (1.Feb.1911)

Evans, W.H.,1949 : A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History).
i-xix + 1-502, pls.1-35. London.

Zhang, J., Cong, Q., Shen, J. & Grishin, N.V.,2022 : Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera).
Insecta Mundi 0921: 1-135.

Xue, G., Qu, P., Li, M., Chiba, H. & Li, W.,2022 :
Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae).
Zootaxa 5205(5): 481–490.

Zhang, J., Cong, Q., Shen, J., Song, L., Opler, P.A. & Grishin, N.V.,2023 :
Additional taxonomic refinements suggested by genomic analysis of butterflies.
Taxon. Rep. Int. Lep. Survey 11(1): 1-25.

♂, Sa Kaeo, Thailand.	♂ + ♀ (mating), Nakhon Nayok, Thailand.
♂, Yala, Thailand.	♂, Yala, Thailand.

1999.09.04 - 2025.01.31.